Checklist of Bolivian Compositae - Royal Botanic Gardens, Kew

Vernacular names: MANZANILLA DEL CAMPO (Cernana & Ariza Espinar, 2008: 7); YATEÍ-KAÁ. (Freire, 1995); MARCELA
BRASILERA, YATEÍ CAÁ (Giangualani, 1976); YATÁ CAÁ, YATEÍ CAÁ ETE (Freire et al., 2006).
Achyrocline alata (Kunth) DC. var. vautheriana (DC.) Baker in Mart., Fl. Bras. 6(3): 117 (1882) = Achyrocline
alata (Kunth) DC.
Achyrocline argentina O. Hoffm., Linnaea 43(2): 135 (1881) = Achyrocline alata (Kunth) DC.
Achyrocline brittoniana Deble & Marchiori, Balduinia 3: 13 (2005) = Achyrocline ramosissima Britton ex
Rusby
*Achyrocline candicans (Kunth) DC., Prodr. 6: 221 (1838) = Achyrocline saturejoides (Lam.) DC.
*Achyrocline celosioides (Kunth) DC., Prodr. 6: 221 (1838).
Gnaphalium celosioides Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 4 (ed. folio): 61 (1818). Type:
‗Crescit juxta Loxam Peruvianorum, alt. 1060 hex. LFloret Julio.‘ Holotype: P-Bonpl.
?Bolivia (La Paz), Ecuador, Peru, Venezuela.
2000–3500 m.
July.
Note: this species was included in Foster‘s list (Foster, 1958) based on a Bang collection (Bang 2022)
attributed to this species by Rusby (1907: 387), but with some doubt.
Achyrocline citrina Griseb., Abh. Königl. Ges. Wiss. Göttingen 24(1): 187 (1879); Symb. Fl. Argent. : 187 (1879)
= Achyrocline flaccida (Weinm.) DC.
Achyrocline flaccida (Weinm.) DC., Prodr. 6: 220 (1838).
Gnaphalium flaccidum Weinm., Flora 3(2): 610 (1820). Type: ‗L. In Brasilia D. Langsdorff.‘ Holotype: possibly LE.
Achyrocline citrina Griseb., Abh. Königl. Ges. Wiss. Göttingen 24(1): 187 (1879); Symb. Fl. Argent. : 187 (1879).
Type: ‗S.: pr. Yacone.‘. Holotype: GOET; isotype: CORD. Note: Anderberg et al. (1996: 3) listed a type
‗Argentina, Salta. Lorentz & Hieronymus s.n., III 1843.‘ in S. Cerana & Ariza Espinar (2008: 8) cited the
CORD isotype as ‚Prov. Salta: Yacone, cerca de Salta, Lorentz et Hieronymus 296, III-1873‘.
Achyrocline saturejoides (Lam.) DC. var. albicans Griseb., Abh. Königl. Ges. Wiss. Göttingen 24(1): 187 (March-
April 1879); Symb. Fl. Argent.: 187 (1879). Type: ‗J.‘ Holotype: GOET.
Argentina, Bolivia (Santa Cruz), Brazil, Paraguay.
Damp margins of thickets and Alnus forest.
0–2000 m.
(October–) March–August.
Note: This species is recorded for Bolivia by Freire & Nieva (1998: 16). The synonymy reflects Giangualani‘s
addition of A. saturejoides var. albicans as a synonym of A. citrina which she recognized as a separate species
(Giangualani, 1976), and Freire (1995) who synonymized them with A. flaccida. Interestingly, Ariza Espinar
(2010: 186) considered Lorentz‘s publication of ‗A. saturejoides var. citrina‘ as the place of publication of a new
entity citing a different collection from that of Grisebach in his Achyrocline citrina. Ariza Espinar considered
Lorentz‘s name a synonym of A. saturejoides.
Vernacular names: MARCELA, VIRA-VIRA (Freire, 1998); FALSO YATEÍ CA‘A, MARCELA, MARCELA HEMBRA, MARCELA MACHO,
VIRA-VIRA (Freire et al., 2006)..
Achyrocline flavescens Griseb., Abh. Königl. Ges. Wiss. Göttingen 19([1]): 181 (1874); Pl. Lorentz.: 133 (1874) =
Achyrocline alata (Kunth) DC.
*Achyrocline hyperchlora S. F. Blake, Bot. Gaz. 74: 415 (1922). Type: ‗BOLIVIA.– Cochabamba, March 14,
1920, E.W.D. and M.M. Holway 406‘. Holotype: US (1058598).
Argentina, Bolivia (Cochabamba), ?Peru.
Humid slopes; c. 3000 m; February–April.
Vernacular name: VIRA-VIRA (Cerana & Ariza Espinar, 2008: 9).
*Achyrocline latifolia Wedd., Chloris Andina 1: 148 (1856). Type: ‗Hab. BOLIVIE: département de Tarija, sur
les berges humides de la côte de Calama, dans la descente de la puna d‘Iscaiachi! (Wedd.).‘ Holotype: P.
Bolivia (Chuquisaca, Santa Cruz, Tarija).
Puna.
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