Checklist of Bolivian Compositae - Royal Botanic Gardens, Kew

Arvet-Touvet, C. (1881). Spicilegium rariorum vel novorum Hieraciorum praecipue americanorum et
europaeorum ... Rigaudin, Grenoble.
Fries, E. (1862). Epicrisis generis Hieraciorum. Edquist & Berglund, Upsaliae. pp. 158.
Schultz-Bipontinus, C. H. (1865). Cichoriacearum Boliviensium novarum a clar. Giliberto Mandon
Pictaviensi lectarum … Linnaea 33(6): 757–758.
Schultz-Bipontinus, C. H. (1865). Pilosellarum boliviensium novarum triga, … Linnaea 33(6): 759–762.
Sleumer, H. (1936). Vermischte Diagnosen. IV. Repert. Spec. Nov. Regni Veg. 41: 117–128. [Compositae: 117–
119].
Sleumer, H. (1956). Die Hieracien Argentiniens unter Berücksichtigung der Nachbarländer. Bot. Jahrb. Syst.
77(1): 85–148.
Zahn, K. H. (1923). Compositae – Hieracium. In: A. Engler (ed.), Das Pflanzenreich, Regni vegetabilis
conspectus. Im auftrage der Preuss. Akademie der Wissenschaften. IV. 280. Abteilung I. Sect. I–XXXIX (Heft
75, 76, 77, 79 & 82). Engelmann, Leipzig. pp. 1–1705.
Key to species (modified and adapted from Sleumer, 1956)
Note: In Sleumer‘s key a number of very specific terms, pertinent to the habit and form of hieracia, were
used. These included:
Aphyllopodous = lacking leaves at flowering time.
Hypophyllopodous = ‗radical leaves present when flowering, but not numerous; used of certain Hieracia‘
(Jackson, 1928).
Phyllopodous = ‗used of the genus Hieracium when the radical leaves are in full vigour‘ at flowering time
(Jackson, 1928).
Subhypophyllopodous = few radical leaves present at flowering time scarcely forming a rosette (possibly a
very loose rosette discernible).
1. Ligule apices markedly ciliate and involucre never or laxly scarcely pilose; leaves usually ±
densely floccose beneath (Sect. Hypocheoridiformia)
H. fimbriatum
Ligule apices ± obviously ciliate and involucre densely pilose to lanate, or ligules never
ciliate; leaves not floccose beneath 2
2. (1) Inflorescence scapiform, rarely 2 or 3 headed, axillary from basal, middle or upper leaves;
involucre large, 12–13 (–14) mm tall, 8–10 mm diam., (densely long-pilose or pilose, never
glandular-pilose) (Sect. ‗Mandonia’)
H. stachyoideum
Inflorescence many- to densely-headed, with fewer from upper leaf axils or usually from
reduced bracts; involucre small to large 3
3. (2) Involucre ± densely long-pilose to villose (lanate), otherwise never obviously long
glandular-pilose, occasionally sparsely, rarely moderately short-glandular, that is short (to
0.5 mm long) glandular-pilose, occasionally sparsely to densely floccose (Sect. Aurelliformia)6
Involucre densely (rarely sparsely) long glandular-pilose, occasionally pilose, occasionally
floccose or involucre floccose, occasionally glandular 4
4. (3) Rosette leaves absent at flowering; inflorescence many- to densely-headed[; involucres small,
subcylindrical, floccose, rarely occasionally laxly glandular-pilose (Sect. Acceipitriniformia)]
H. microcehpalum
Rosette leaves present and green at flowering or present and few; inflorescence few- to
many-headed 5
5. (4) Involucre never or moderately glandular-pilose, always ± densely floccose; style ± yellow
(rosette leaves always present) (Sect. Piloselliformia) 14
Involucre densely to very densely glandular-pilose, otherwise ± densely pilose, otherwise
floccose; style usually dark (rosette leaves present or relatively few) (Sect. Adenothyrsa) 16
6 (3). Leaves distinctly rosettiform in flower; cauline leaves 1–3, rapidly falling, lower forming a
rosette 7
Basal leaves few, scarcely forming a rosette when plant in flower; cauline leaves 3–6–8 (12,
rarely to 15), gradually falling 11
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