Checklist of Bolivian Compositae - Royal Botanic Gardens, Kew

Capitula few, laxly cymose-corymbose; leaves subulate, margins entire or lobate S. subulatus
Several species also key out into the Ser. Suffruticosi key below!
Key to species of Ser. Graveolentes
Shrublets with divaricate spiny branches
Spineless shrubs
Key to species of Ser. Suffruticosae
Species marked ‗*‘ are also keyed out here as well as in the Ser. Corymbosi key!
S. spinosus
S. nutans
1. Capitula heterogamous, marginal florets female, rayed or shortly tubular; disc florets
hermaphrodite, tubular 2
Capitula homogamous, florets all hermaphrodite, tubular 7
2. (1) Plants densely tomentose 3
Plants glabrescent 5
3. (2) Involucre 6–7 mm tall, 6 mm diam. S. viridilacus
Involucre 12–15 mm tall, c. 20 mm diam. 4
4. (3) Leaves 3–9 cm long, oblong; ray limbs scarcely excedding involucre S. octophyllus
Leaves 1–3 cm long, ovate-lanceolate; ray limbs well exceeding involucre S. hohenackeri
5. (2) Involucre 5–6 mm tall; leaves linear, margins entire or dentate S. potosianus
Involucre 10 mm tall 6
6. (5) Leaves 5–10 mm long, with 2–3 pairs of triangular segments 1–3 mm long *S. viridis
Leaves 10–60 mm long with 1–3 pairs of linear segments 2–20 mm long *S. subulatus
7. (1) Plants glandular-pubescent 8
Plants tomentose, lanose or glabrous 10
8. (7) Achenes densely pubescent; leaves 1–2 cm long, margins profoundly dentate or lobate
S. adenophyllus
Achenes glabrous; leaves 1–3.5 cm long, margins entire (rarely subdentate) 9
9. (8) Leaves 1–5 mm wide; calycular bracts oblong, reaching almost the middle of the involucre
*S. rufescens
Leaves 4–10 mm wide; calycular bracts narrowly lanceolate almost as long as the involucre
*S. wedglacialis
10. (7) Leaves narrowly linear, apices very acute, margins entire or with a few linear lobes 11
Leaves linear-lanceolate, linear-spathulate, elliptic, obovate, ovate or circular, apices
rounded or obtuse, rarely acute, margins entire, crenate or pinnatisect 14
11. (10) Phyllaries 10–18 12
Phyllaries 5–7 13
12. (11) Leaves tomentulose, 10–30 mm long; phyllaries 10–15; capitula relatively few at stem apices
*S. chrysolepis
Leaves 25–80 mm long; phyllaries 13–18; capitula solitary
S. scorzonerifolius
13. (11) Involucre 4–5 mm tall; leaves 10–15 mm long S. helianthemoides
Involucre 9–10 mm tall; leaves 50 mm long
*S. quinquelepis
14. (10) Involucre 11–15 tall; phyllaries 16–25 15
Involucre shorter 16
15. (14) Leaves 15–25 mm long, margins entire or with 2–3 pairs of large teeth or lobes, laxly lanose
S. aquilaris
Leaves 40–120 mm long; margins profoundly pinnatisect, with 5–12 pairs of linear segments,
glabrous
S. jarae
16. (14) Capitula small, sessile; involucre 5 mm tall 17
Capitula larger 18
17. (16) Phyllaries 8; leaves entire, glabrous S. humillimus
Phyllaries 13; leaves entire or dentate, glabrous or lanuginose
S. vegetus
18. (16) Plants tomentose, with the exception of underside of leaves 19
Plants glabrous 23
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