Checklist of Bolivian Compositae - Royal Botanic Gardens, Kew

18. (6) Style arms long-clavate, thickened in both width and thickness, few exceptions with 10-
ribbed achenes, plumose pappus or those with peg-like setulae on achenes; upper corolla
limb often constricted beneath corolla lobes (subtribe Alomiinae) 19
Style arms not thickened or thickened only at tips, if long-clavate then flattened; achenes
never 10-ribbed, never with plumose pappus setae and never with peg-like eglandular
achene setulae; corolla limb cylindrical or funnelform beneath corolla lobes 21
19. (18) Style base with distinctly enlarged basal node; corollas glabrous outside 20
Style base lacking enlarged basal node; corollas glandular-punctate outside Helogyne
20. (19) Achenes 8–10-ribbed; annual herb Brickellia diffusa
Achenes 5-ribbed; erect or spreading to arching subshrubs or shrubs Austrobrickellia
21. (18) Style base glabrous and basal node absent; inner phyllaries usually deciduous 22
Style base with basal node or pubescent or both 33
22. (21) Capitula always with 5 florets; style arm apices papillose; pappus setae sometimes with
bulbous-tipped apical cells; plants of eastern South America (subtribe Disynaphiinae) 23
Capitula with 1–300 florets, rarely consistently 5-flowered; style arms usually smooth at least
at apices; pappus setae never with bulbous-tipped apical cells; plants throughout tropical
and subtropical America 25
23. (22) Leaves pinnately or bipinnately lobed; style arm appendages with long narrow papillae
Acanthostyles
Leaves simple; style arms appendages short-papillose 24
24. (23) Inflorescences densely paniculate Raulinoreitzia
Inflorescences densely corymbose
Campovassouria
25. (22) Receptacles glabrous; anther collar usually less than 5 times as long as wide (subtribe
Critoniinae) 26
Receptacles usually pubescent; anther collar usually more than 5 times as long as wide
(subtribe Hebecliniinae)
Hebeclinium
26. (25) Leaves distinctly alternate Bishovia
Leaves mostly opposite or subopposite 27
27. (26) Phyllaries distant or weakly subimbricate in up to 3 rather irregular series 28
phyllaries subimbricate or imbricate in 4 or more gradate series, inner phyllaries sometimes
easily deciduous 30
28. (27) Apical anther appendages absent, often appearing as two minute lobes; phyllaries distant
Ophryosporus
Apical anther appendages distinct, 1/3 as long as wide or longer; phyllaries weakly
subimbircate 29
29. (28) Capitula in dense spherical clusters Sphaereupatorium
Capitula not in dense spherical clusters
Koanophyllon
30. (27) Apical anther appendages shorter than wide, truncate to bilobed; style arms often with
abruptly enlarged apices
Koanophyllon
Apical anther appendages about as long as wide or longer, usually with rounded apices,
rarely retuse; style arms with abruptly enlarged apices in some Critonia 31
31. (30) Leaves with translucent or lens-like secretory pockets when viewed against light, lacking
glandular-punctae; coarse vines and shrubs
Critonia
Leaves lacking secretory pockets, often with glandular-punctae 32
32. (31) Pappus setae with markedly dilated spinose apices; capitula with 50–70 florets Amboroa
Pappus setae without dilated or spinose apices; capitula with 10–12 florets Lorentzianthus
33. (21) Style base lacking basal node, pubescent; capitula with less than 20 florets; receptacles
epaleaceous; apical cells of pappus setae usually rounded (subtribe Eupatoriinae) 34
Style base inflated above nectary, glabrous or pubescent; capitula usually with more than 20
florets, sometimes receptacle paleaceous; apical cells of pappus setae acute (subtribe
Ayapaninae) 36
34. (33) Achene body setuliferous 35
Achene body only with glands
Austroeupatorium
35. (34) Inflorescences pyramidal or thyrsoid Stomatanthes
Inflorescences corymbose
Hatschbachiella
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