Checklist of Bolivian Compositae - Royal Botanic Gardens, Kew

7. (6) Inflorescence laxly ± markedly branched or continuously to median leaves (rarely markedly)
branches remote, 2–5, slender, 1 or 2 (–3)-headed; [leaves densely whitish-cinereous-villose
or -lanate on both surfaces; involucre short-cinereous-lanate and densely floccose
(Cochabamba)]
H. herzogianum
Inflorescence ± compact, branches short, 1–3 headed 8
8. (7) Involucre clearly (sparsely or densely) short-glandular 9
Involucre never or rarely laxely short-glandular 10
9. (8) Involucre subovate-cylindrical, 7–8 mm c. 5 mm diam., ± pilose throughout; delicate,
subscapose, 6–12 (–15) cm tall, ± densely pilose (Cochabamba, La Paz) H. mandonii
Involucre ovate, 8–9 mm × 6–7 mm diam., blackish-tangled villose in lower half, long-pilose
above; stem 15–25 cm tall (rarely shorter), usually sublanate-short pilose (Cochabamba, La
Paz)
H. trichodontum
10. (8) Involucre narrowly ovate-cylindrical, ± densely floccose, 6–7 mm tall
H. argentinense (delicate plants)
Involucre ovate-cylindrical, sparsely or scarcely floccose at base, 8–10 mm tall (Chuquisaca)
H. eriocephalum
11. (6) Few radical leaves present at flowering time, scarcely rosettiform; stems slender or robust;
cauline leaves not very or strongly clasping
H. argentinense
Radical leaves distinctly rosettiform at flowering time; stems robust; cauline leaves markedly
amplexicaul, 8–12 (–15), few or none remaining at flowering 12
12. (11) Leaves oblong-ovate, subobtuse, 5–6(–7) cm long; involucres 7–8 mm tall, moderately
blackish-pilose; stem hairs dense, setiform, patent (5–6 mm long) 13
Leaves (subobovate) oblong to elongate-oblong, acuminate, (8) 10 to 15 cm long; involucre 8–
10 (–12) mm tall, densely long blackish-pilose (Cochabamba, La Paz)
H. tacense (? incl. H. trollii)
13. (12) Phyllaries densely short-glandular pubescent beneath pilose hairs, dense at base and
floccose towards apex (Chuquisaca)
H. boliviense
Phyllaries sparsely short-glandular beneath pilose hairs, floccose almost throughout (Tarija)
H. padcayense
14. (5) Involucre laxly or moderately glandular-pilose, densely floccose; involucre small, 4–5 mm
tall (La Paz)
H. apoloense
Involucre ± densely glandular-pilose, scarcely to moderately floccose 15
15. (14) Involucre 6–8 mm tall; inflorescence many-headed, subcorymbose-paniculate (Chuquisaca,
Tarija)
H. fulvipes
Involucres (8–) 9 (–10) mm tall; inflorescence few-headed, ± corymbose, branches markely
forked and single-headed (La Paz)
H. mapirense
16. (5) Involucre ± black glandular-pilose, rarely subsetulose, rarely subfloccose towards base (Sect.
Adenothyrsa s.s.) 17
Involucre black-glandular-pilose and also distinctly pilose (setose), frequently somewhat
floccose or black-glandular-pilose and also ± densely floccose or never pilose, or black
(rarely yellowish)- glandular-pilose and also pilose (setose) and floccose 20
17. (16) Inflorescence branches (to 8) short, ± erect few-headed, lowermost remotely thyrsoid or
subpaniculate; radical leaves present at flowering time; cauline leaves smaller; (stems often
softly patent setose-pilose) 18
Inflorescence branches ± elongate and thyrsoid towards apex, paniculate below; cauline
leaves moderately abruptly smaller 19
18. (17) Stem base densely setose-pilose, upwards laxly setose-pilose; involucre 6–7.5 mm tall;
cauline leaves to 12
H. dasychaetocomum
Stem ± patent setose-pilose throughout; involucre (9–) 10–11 mm tall; cauline leaves 5–7
H. asplundii
19. (17) Involucre 4–5 mm tall; inflorescence diffuse or strongly intermediate, with 6–8 terminal
branches, remaining strongly remote and long-squarrose and corymbiform with many
heads; rosette leaves and stem base floccose; cauline leaves 3–4, remote, ± oblong, never or
hardly amplexicaul
H. hauthalianum
Involucre 6– (rarely –9) mm, subfloccose beneath; inflorescence thyrsoid-paniculate, many-
(to 35)-headed, branches ± elongate arcuate-ascending with final subracemose-branching;