Checklist of Bolivian Compositae - Royal Botanic Gardens, Kew

Trixis divaricata (Kunth) Spreng. var. sprengeliana (Gardner) Baker in Mart., Fl. Bras. 6(3): 385 (1884) = Trixis
divaricata (Kunth) Spreng.
Trixis flexuosa (Kunth) Spreng., Syst. Nat. 3: 501 (1826) = Trixis divaricata (Kunth) Spreng.
Trixis frutescens P.Browne var. cacalioides Griseb., Abh. Königl. Ges. Wiss. Göttingen 19([1]): 199 (1874); Pl.
Lorentz. 19: 151 (1874) = Trixis grisebachii Kuntze
Trixis frutescens R.Browne var. denticulata Less., Syn. Gen. Comp. : 414 (1832), nom. illegit. based on Perdicium
cacalioides Kunth = Trixis cacalioides (Kunth) D. Don
Trixis grisebachii Kuntze, Revis. Gen. Pl. 3(3): 183 (1898). Types: ‗Argentina: Jujuy (Lorentz & Hieronymus),
im Berliner bot. Museum ausserdem von Tucuman: Cuesta de Juntas (Lorentz). Note: it is clear that the
Lorentz specimen was in B, presumed destroyed and Wetter & Zanoni (1985: 339) noted a Lorentz &
Hieronymus s.n. type in NY (as ‗Argentina. Jujuy, Apr 1872‘). These are taken as the syntypes. Katinas (1996:
64) merely noted the NY type, but cited a ‗holotype‘ for ‗Trixis frutescens P. Browne var. cacalioides Griseb.‘ (as
a synonym) as ‗Argentina, Prov. Tucumán. Sierra de Tucumán, Cuesta v. Juntas, Lorentz 324‘ in B, based on
photograph in F (16098). The lectotype is probably best chosen as Lorentz & Hieronymus s.n., NY (00274292).
Argentina, Bolivia (Tarija).
Roadsides, field margins, railways.
0–2500 m.
December–August.
Vernacular name/s: TIAN-TIAN; TIEN-TIEN (Katinas (1996: 66).
Trixis (§ Aplochlaenae) hexantha S. Moore, J. Bot. 52: 264 (1914) = Trixis cacalioides (Kunth) D. Don
Trixis hoffmanniana Taub., Bot. Jahrb. Syst. 21(4): 456 (1896) = Trixis ophiorhiza Gardner ssp. ophiorhiza
Trixis neaeana DC., Prodr. 7: 67 (1838) = Trixis cacalioides (Kunth) D. Don
Trixis odoratissima Gardner, London J. Bot. 6: 461 (1847) = Trixis divaricata (Kunth) Spreng.
Trixis ophiorhiza Gardner, London J. Bot. 6: 461 (1847). Types: ‗HAB. Between Capella da Passe and San
Pedro, Province of Goyaz ([Gardner] 4266), and between the Rio de San Francisco and Formigas, Province of
Minas Geraes ([Gardner] 4959). May and July 1840.‘ Syntypes: K. Lectotype (selected by Katinas, 1996: 78, but
cited as ‗holotype‘): Gardner 4959, BM; isolectotypes: B, GH, K, NY (00274297, 00274299). Isosyntype (Gardner
4266): K, NY (00274298).
ssp. ophiorhiza
Trixis chionopappa Sch.Bip. in Mart. Fl. Bras. 6(3): 386 (1884), nom. nud. pro syn.
Trixis hoffmanniana Taub., Bot. Jahrb. Syst. 21(4): 456 (1896). Type: ‗Habitat in locis Cerrados vocatis in
fluminis Paranahyba ditione: ULE n. 2969. – Flor. m. Jul.‘ Holotype: ?; isotype: HBG.
Trixis cerroleonensis Soria & Zardini, Ann. Missouri Bot. Gard. 78(2): 531 (1991). Type: ‗Paraguay. Chaco:
Cerro Léon, 8 Aug. 1983, Hahn 1555‘. Holotype: PY; isotypes: LP, MO, SI.
Brazil, Bolivia (Santa Cruz), Paraguay.
Cerrado, disturbed soils.
To 1000 m.
June–September.
Vernacular name: MBÓI POHÃ (Cabrera, 1998).
Trixis papillosa Gillies ex D. Don, Phil. Mag. 2: 388 (1832) = Trixis cacalioides (Kunth) D. Don
Trixis paradoxa Cass., Opusc. Phytol. 2: 160 (1826) = Trixis cacalioides (Kunth) D. Don
Trixis ravenii Zardini & Soria, Ann. Missouri Bot. Gard. 78(2): 531 (1991) = Trixis spicata Gardner
*Trixis rigida J. Koster, Blumea 5(3): 682 (1945) = Trixis aggregata Rusby
Trixis salicifolia D. Don, Trans. Linn. Soc. London 16(2): 298 (1830) = Trixis divaricata (Kunth) Spreng.
Trixis spicata Gardner, London J. Bot. 6: 462 (1847). Type: ‗HAB. Grassy campos near the Rio Claro, Province
of Minas Geraes. [Sept. 1840]‘ [Gardner] 4957. Note: Katinas (1996: 91) noted the ‗Holotype: BM; isotype: K ...‘
Trixis ravenii Zardini & Soria, Ann. Missouri Bot. Gard. 78(2): 531 (1991). Type: ‗Paraguay, Amambay: Arroyo
Estrella, 2220'S, 5620'W, 7 May 1989, Soria, Bacigalupo & Fortunato 3825‘. Holotype: FCQ.
Bolivia (Santa Cruz), Brazil, Paraguay.
Grassland.
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