Checklist of Bolivian Compositae - Royal Botanic Gardens, Kew

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*Achyrocline polycephala Rusby, Bull. New York Bot. Gard. 4(14): 388 (1907) = Achyrocline tomentosa Rusby Achyrocline pterocaula DC., Prodr. 6: 221 (1838) = Achyrocline alata (Kunth) DC. *Achyrocline ramosissima Britton ex Rusby, Mem. Torrey Bot. Club 3(3): 57 (1893). Type: [Bolivia:] ‗Vic. Cochabamba, 1891 ([Bang] 99).‘ Holotype: NY (00169516); isotype: NY (00169515). Note: Rusby (1893) referred to the place at which Britton proposed a combination (in Achyrocline) based on Schultz Bipontinus‘ nom. nud., Gnaphalium (Achyrocline) ramosissimum (based on Mandon 157). Deble & Marchiori‘s new name, q.v. Achyrocline brittoniana, is unnecessary. Gnaphalium ramosissimum Sch.Bip., Bonplandia 4(4): 54 (1856), nom. nud., non Nutt. (1848) (= Pseudognaphalium ramosissimum (Nutt.) Anderb.) Gnaphalium (Achyrocline) ramosissimum Sch.Bip., Bull. Soc. Bot. France 12: 80 (1865); Linnaea 34(5): 532 (Feb 1866), nom. nud. (based on Mandon 157). Achyrocline brittoniana Deble & Marchiori, Balduinia 3: 13 (2005). Note: Deble & Marchiori (2005: 13), followed by Cerana & Ariza Espinar (2008: 8), took Rusby‘s publication of the name Achyrocline ramosissima to be a combination (as it appeared in Rusby, 1893: 57). Argentina, Bolivia (Cochabamba, Santa Cruz, Tarija), Peru. Puna, amongst rocks. 2200–4500 m September–December. *Achyrocline rufescens (Kunth) DC., Prodr. 6: 220 (1838) = Achyrocline alata (Kunth) DC. Achyrocline rupestris Cabrera, Darwiniana 9(1): 43 (1949) = Achyrocline tomentosa Rusby *Achyrocline saturejoides (Lam.) DC., Prodr. 6: 220 (1838). Gnaphalium saturejoides Lam., Encycl. 2: 747 (1786). Type: ‗Commerson a trouvé cette plante à Monte Video. L. (v.s.).‘ Holotype: P-LA (323/12). Gnaphalium candicans Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Sp. 4 (ed. folio): 62 (1818). Type: [Ecuador:] ‗Crescit in Regno Quitensi inter pagum Ticsan et urbem Alausi, alt. 1250 hex. KFloret Julio.‘ [Humboldt & Bonpland ‗n. 3244. Alausi‘]. Holotype: P-Bonpl. Achyrocline vargasiana DC., Prodr. 6: 220 (1838). Type: [Venezuela:] ‗Lcircà Caracas legit cl. Jos. Vargas. ... (v.s.)‘. Holotype: G-DC. *Achyrocline candicans (Kunth) DC., Prodr. 6: 221 (1838). *Gnaphalium satureioides Lam. var. candicans (Kunth) Kuntze, Revis. Gen. Pl. 3(3): 153 (1898). Argentina, Bolivia (Chuquisaca, Cochabamba, La Paz, Santa Cruz, Tarija), Brazil, Colombia, Ecuador, Paraguay, Peru, Uruguay, Venzuela. Tracksides, amongst rocks, marshes, damp flushes, grassland. 0–2500 m October–August. Chuquisaca: Wood 8290, 11.4.1994 (K). Vernacular names: MARCELA; VIRÁ-VIRÁ GUAZÚ; VIRA VIRA (Cerana & Ariza Espinar, 2008: 10); ALQUITRÁN (Freire, 1995); MARCELA MANZANILLA (Freire, 1998); ALQUITRÁN, MARCELA, MARCELA DEL CAMPO, MARCELA HEMBRA, MARCELA MANZANILLA, MARCELITA, PAG(A)RÁ LAURÓ, PLANTA DE LA ABEJA, VIRÁ-VIRÁ GUAZÚ, YATEÍ CA‘Á, YATEÍ CA‘A, YATEÍ CAÁ ESTERO (Freire et al., 2006). Achyrocline saturejoides (Lam.) DC. var. albicans Griseb., Abh. Königl. Ges. Wiss. Göttingen 24(1): 187 (March- April 1879); Symb. Fl. Argent.: 187 (1879) = Achyrocline flaccida (Weinm.) DC. *Achyrocline tomentosa Rusby, Bull. New York Bot. Gard. 4(14): 388 (1907). Type: [Bolivia:] ‗ ‖A small plant, a few feet high, with white flowers; rather scarce in very dry soil on rocks.‖ Coripata, April 10, 1894. ([Bang] No. 2124.)‘ Holotype: NY (00149892); isotype: CORD, NY (00149893), US(325879). Achyrocline polycephala Rusby, Bull. New York Bot. Gard. 4(14): 388 (1907). Type: [Bolivia:] ‗ ‖In dry gravelly soil, climbing 5 or 6 feet, the flowers white.‖ Coroico, July 11, 1894. ([Bang] No. 2336.)‘ Holotype: NY (00149889); isotypes: F (163885), K, NY (00149888 – ex College of Pharmacy Herbarium), US (00032932). Achyrocline rupestris Cabrera, Darwiniana 9(1): 43 (1949). Type: ‗Argentina. – Salta: Dep. San Antonio de los Cobres, Quebrada de Polvorilla, entre rocas a 4300 m s. m., leg. A. L. Cabrera, 8395, 2-II-1944‘. Holotype: LP. Argentina, Bolivia (Santa Cruz), Paraguay, ?Peru. Amongst rocks, Puna.
500–3500 (–4500) m. March–April. Note: Cerana & Ariza Espinar (2008: 10 & 12) clearly regarded A. rupestris and A. tomentosa as distinct species, based primarily upon plant size; this view is not maintained here. Achyrocline vargasiana DC., Prodr. 6: 220 (1838) = Achyrocline saturejoides (Lam.) DC. *Achyrocline vauthieriana DC., Prodr. 6: 220 (1838) = Achyrocline alata (Kunth) DC. *Achyrocline venosa Rusby, Mem. Torrey Bot. Club 3(3): 57 (1893). Type: [Bolivia:] ‗Vic. Cochabamba, 1891. Specimen unique, sent as part of [Bang] 99. May be called [Bang] 99a. At first regarded as Gnaphalium.‘ Holotype: NY (149894). Note: This specimen was determined by Schofield (1972) as an isotype; this is unlikely since Rusby‘s comment was quite specific ‗Specimen unique, ...‘ Argentina, Bolivia (Cochabamba), Paraguay, Venezuela. Distribution provided in Freire & Nieva (1998: 19). Sandy soils, grassland, tracksides. 1500–4000 m. September–June. Vernacular names: VIRA-VIRA CHACO, VIRA-VIRA GUASU (Freire, 1998). Achyropappus Link & Otto, Ic. Pl. Rar. Pl. 30 (1829), non Achyropappus Kunth (1818) = Schkuhria Roth Achyropappus neo-mexicana (A. Gray) Rydb., Fl. Colorado : 377 (1906) = Schkuhria multiflora Hook. & Arn. Achyropappus schkuhrioides D. Don ex Hook. & Arn., non A. schkurioides Link & Otto (1829) = Schkuhria multiflora Hook. & Arn. Achyrophorus Adans., Fam.. Pl. 2: 112 (1763) = Hypochaeris L. Achyrophorus Adans. sect. II. Oreophila (D. Don) DC., Prodr. 7: 93 (1838) = Hypochaeris L. Achyrophorus acaulis Remy in Gay, Fl. Chil. 3: 448 (late 1847 or early 1849) = Hypochaeris acaulis (Remy) Britton Achyrophorus albiflorus Sch.Bip., Jahresber. Pollichia 16/17: 52 (1859) = Hypochaeris sessiliflora Kunth Achyrophorus brasiliensis (Less.) Sch.Bip., Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 21: 106 (1845) = Hypochaeris chillensis (Kunth) Hieron. Achyrophorus chilensis Sch.Bip., Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 21: 104 (1845) = Hypochaeris chillensis (Kunth) Hieron. Achyrophorus chillensis (Kunth) Sch.Bip., Jahresber. Pollichia 16/17: 57 (1859) = Hypochaeris chillensis (Kunth) Hieron. Achyrophorus cryptocephalus Sch.Bip. Bonplandia 4(4): 54 (1856) = Hypochaeris eriolaena (Sch.Bip.) Reiche Achyrophorus elatus Wedd., Chloris Andina 1: 220 (1857) = Hypochaeris elata (Wedd.) Griseb. Achyrophorus eriolaenus Sch.Bip., Jahresber. Pollichia 16/17: 47 (1859) = Hypochaeris eriolaena (Sch.Bip.) Reiche Achyrophorus hoehenackeri Sch. Bip., Bonplandia 4(4): 54 (1856) = Hypochaeris hohenackeri (Sch.Bip.) Domke Achyrophorus humboltdii Sch.Bip., Jahresber. Pollichia 16/17: 52 (1859) = Hypochaeris sessiliflora Kunth Achyrophorus meyenianus (Meyen & Walp.) Walp., Rep. 6: 336 (1846-47) = Hypochaeris meyeniana (Meyen & Walp.) Griseb. Achyrophorus meyenianus Walp. var. ciliata Wedd., Chloris Andina 1: 220 (1857) = Hypochaeris sessiliflora Kunth Achyrophorus quitensis Sch.Bip., Nov. Actorum Caes. Leop.-Carol. Nat. Cur. 21: 100 (1845), nom. nud. = Hypochaeris sessiliflora Kunth Achyrophorus quitensis Sch.Bip. ex Wedd., Chloris Andina 1: 219 (1957), nom. illegit. superfl. = Hypochaeris sessiliflora Kunth Achyrophorus quitensis Sch.Bip. var. sonchoides (Kunth) Wedd., Chloris Andina 1: 219 (1857) = Hypochaeris sessiliflora Kunth Achyrophorus sagittatus Phil., Anales Univ. Chile 36: 178 (1870) = Hypochaeris chillensis (Kunth) Hieron. Achyrophorus sessiliflorus (Kunth) DC., Prodr. 7: 95 (1838) = Hypochaeris sessiliflora Kunth Achyrophorus setosus Wedd., Chloris Andina 1: 220 (1857) = Hypochaeris echegarayi Hieron. Achyrophorus stenocephalus A. Gray ex Wedd., Chloris Andina 1: 221 (1857), nom. nud. = Hypochaeris stenocephala Kuntze 35
Page 1 and 2: AN ANNOTATED PRELIMINARY CHECKLIST
Page 3 and 4: determination of J. R. I. Wood et a
Page 5 and 6: The Checklist Key to tribes of Comp
Page 7 and 8: 12. (7) Appendages of fertile disc
Page 9 and 10: 18. (6) Style arms long-clavate, th
Page 11 and 12: 15. (14) Female floret corollas tub
Page 13 and 14: 51. (50) Plants mainly prostrate; c
Page 15 and 16: 3. (2) Achenes long-beaked; corolla
Page 17 and 18: glabrous; pappus setae usually unis
Page 19 and 20: 7. (4) Florets 2-4 per capitulum; c
Page 21 and 22: Noticastrum DC. 4 Novenia S. E. Fre
Page 23 and 24: Ambrosia L. 3 Aphanactis Wedd. 1 1
Page 25 and 26: Gamochaeta Wedd. 8 Gnaphalium L. 3
Page 27 and 28: Werneria Kunth 12 Xenophyllum V. A.
Page 29 and 30: A ?Abasoloa La Llave & Lex., Nov. V
Page 31 and 32: Sauce Chico (LOR[ENTZ]. u. NIEDERL[
Page 33: Vernacular names: MANZANILLA DEL CA
Page 37 and 38: Spilanthes arrayana Gardner, London
Page 39 and 40: Ceratocephalus americanus (L.f.) Ku
Page 41 and 42: Adenostemma gymnocoronis Sch.Bip. e
Page 43 and 44: Ageratina acevedoi H. Rob., Phytolo
Page 45 and 46: Eupatorium tenue R. E. Fr., Arkiv B
Page 47 and 48: Vernacular names: CAMARÁ-OPELA, CA
Page 49 and 50: *Franseria recurva Rusby, Bull. New
Page 51 and 52: Angelphytum herzogii (Hassl.) Prusk
Page 53 and 54: Molfino, J. F. (1953). Una nueva es
Page 55 and 56: Argentina, Bolivia (Chuquisaca, San
Page 57 and 58: Aster marginatus Kunth var. argent
Page 59 and 60: Austroeupatorium inulifolium (Kunth
Page 61 and 62: Eupatorium hasslerianum Chodat, Bul
Page 63 and 64: ?Ayapanopsis tucumanensis (Lillo &
Page 65 and 66: Baccharis L. sect. Macrophyllae Hee
Page 67 and 68: Müller, J. (2006). Systematics of
Page 69 and 70: Pingraea articulata (Lam.) F. H. He
Page 71 and 72: October-December. *Baccharis caprar
Page 73 and 74: *Baccharis conwayi Rusby, Bull. New
Page 75 and 76: Molina venosa Ruiz & Pav., Syst. Ve
Page 77 and 78: also under B. medullosa. Teodoro Lu
Page 79 and 80: *Baccharis floribunda Kunth in Humb
Page 81 and 82: Baccharis cordobensis Heering, Jahr
Page 83 and 84: *Baccharis mapirensis Rusby, Mem. T
Page 85 and 86:
Baccharis brachylaenoides DC., Prod
Page 87 and 88:
2800-3700 m. July-August. Vernacula
Page 89 and 90:
*Baccharis condensata Rusby, Descr.
Page 91 and 92:
*Baccharis mapirensis Rusby, Mem. T
Page 93 and 94:
Southwestern USA south to Patagonia
Page 95 and 96:
*Baccharis subalata Wedd., Chloris
Page 97 and 98:
ssp. tola var. tola Chile, Bolivia
Page 99 and 100:
unnamed, unranked ‗Wood-sides of
Page 101 and 102:
Robinson, H. (2006). Baltimora. In:
Page 103 and 104:
February-November. Barnadesia corym
Page 105 and 106:
Barnadesia woodii D. J. N. Hind, Ke
Page 107 and 108:
Type: Belloa chilensis (Hook. & Arn
Page 109 and 110:
Gnaphalium piptolepis (Wedd.) Grise
Page 111 and 112:
open, grassy páramo, alt. circ. 37
Page 113 and 114:
label ‗Bidens odorata/Icon. tab.
Page 115 and 116:
Bonpland ‗mss. n. 698. Caracas‘
Page 117 and 118:
Coreopsis trifoliata Bertol., Fl. G
Page 119 and 120:
October-April. Vernacular names: AM
Page 121 and 122:
Bigelowia DC. sect. Diplostephioide
Page 123 and 124:
Blumea DC., Arch. Bot. 2: 514 (1833
Page 125 and 126:
C Cacalia L., Sp. Pl. : 753 (1753)
Page 127 and 128:
Calea L. subgen. Haplocalea Less.,
Page 129 and 130:
Cerrado, grassland. 290-1500 m. Feb
Page 131 and 132:
Calliopsis Rchb., Icon. Descr. Pl.
Page 133 and 134:
Vernacular names: CHARRÚA KA‘A,
Page 135 and 136:
Carphobolus Schott subgen. Eucarpho
Page 137 and 138:
Centratherum punctatum Cass., Dict.
Page 139 and 140:
Centrospermum Kunth in Humb., Bonpl
Page 141 and 142:
Bolivia (?La Paz), Peru. Puna Perua
Page 143 and 144:
Chaptalia diversifolia Greene, Leaf
Page 145 and 146:
Note: The combination ‗Leria nuta
Page 147 and 148:
Cordillère de Sorata!, sur le chem
Page 149 and 150:
Chevreulia stolonifera Cass., Dict.
Page 151 and 152:
Chromolaena arnottiana (Griseb.) R.
Page 153 and 154:
Eupatorium concinnum Hook. & Arn.,
Page 155 and 156:
Vernacular names: SANATODO (Cabrera
Page 157 and 158:
Eupatorium ramosissimum Gardner, Lo
Page 159 and 160:
*Chrysanthellum americanum (L.) Vat
Page 161 and 162:
Key to species 1. Leaf lamina ovate
Page 163 and 164:
Flowering throughout the year. Vern
Page 165 and 166:
Lectotype: C. heterophyllum (L.) J.
Page 167 and 168:
Cirsium lanceolatum (L.) Scop. ssp.
Page 169 and 170:
Clibadium micranthum O. E. Schulz,
Page 171 and 172:
Cnicus strigosus Hoffmanns. & Link,
Page 173 and 174:
limits of C. bonariensis and C. flo
Page 175 and 176:
Conyza deserticola Phil., Anales Mu
Page 177 and 178:
Conyza pulchella Kunth in Humb., Bo
Page 179 and 180:
León above Oña. Alt. ca. 3000 m.,
Page 181 and 182:
typewritten label with the same tex
Page 183 and 184:
collection as ‗H. A. Weddell 3663
Page 185 and 186:
Key to species 1. Receptacle pilose
Page 187 and 188:
Cotula cabrerae Caro, Kurtziana 1:
Page 189 and 190:
Vernonia cajamarcensis H. Rob., Phy
Page 191 and 192:
as C. humile might also key out to
Page 193 and 194:
*Culcitium neaei Sch.Bip. ex Wedd.,
Page 195 and 196:
5. (3) Achenes glabrous; leaves 1-2
Page 197 and 198:
Dasyphyllum ferox (Wedd.) Cabrera,
Page 199 and 200:
Delprete, P. G. (1994). Systematic
Page 201 and 202:
yellow flowers, scarce in forest-mo
Page 203 and 204:
Note: ‗Senecio curvidens Sch.Bip.
Page 205 and 206:
2000-2200 m. August-September. Desm
Page 207 and 208:
Note: This species apparently grows
Page 209 and 210:
Bolivia (?), Chile, Peru. Altiplano
Page 211 and 212:
E Echetrosis Phil., Anal. Univ. Chi
Page 213 and 214:
huc reitè referencdae? E. erecta L
Page 215 and 216:
Platystephium graveolens Gardner, L
Page 217 and 218:
Elephantopus crispus (Cass.) D. Die
Page 219 and 220:
Chaetospira funckii (Turcz.) S. F.
Page 221 and 222:
Xyridopsis Welw. ex B. Nord., Opera
Page 223 and 224:
Wahlenbergia Schumach., Beskr. Guin
Page 225 and 226:
Enydra [sub Enhydra] rivularis Gard
Page 227 and 228:
*Erechtites hieraciifolia (L.) Raf.
Page 229 and 230:
Note: Nordenstam & Lut (2009) sugge
Page 231 and 232:
Phalacroloma Cass., Dict. Sci. Nat.
Page 233 and 234:
Erigeron (Eschenbachia) niveus Sch.
Page 235 and 236:
*Eupatorium amygdalinum Lam., Encyc
Page 237 and 238:
*Eupatorium conyzoides Vahl var. ci
Page 239 and 240:
Eupatorium heptanthum Sch.Bip., Bon
Page 241 and 242:
*Eupatorium longipetiolatum Sch.Bip
Page 243 and 244:
Eupatorium piquerioides DC., Prodr.
Page 245 and 246:
Eupatorium stigmatosum Chodat var.
Page 247 and 248:
F Fabera Sch.Bip., Nov. Actorum Aca
Page 249 and 250:
Facelis retusa (Lam.) Sch.Bip. var.
Page 251 and 252:
Fleischmannia bridgesii (B. L. Rob.
Page 253 and 254:
Flotovia divaricata (Griseb.) Hiero
Page 255 and 256:
G Gaertneria Medik., Phil. Bot. 1:
Page 257 and 258:
*Galinsoga calva Rusby, Mem. Torrey
Page 259 and 260:
Sabazia urticifolia (Kunth) DC., Pr
Page 261 and 262:
Cabrera, A. L. (1961). Observacione
Page 263 and 264:
Argentina, Bolivia (Chuquisaca), Ch
Page 265 and 266:
*Gnaphalium ecuadoriense Hieron. va
Page 267 and 268:
Moquinia DC., Prodr. 7: 22 (1838),
Page 269 and 270:
Spadonia polymorpha Less. [var.] e
Page 271 and 272:
Brachyris mandonii Sch.Bip., Linnae
Page 273 and 274:
Gymnostyles stolonifera (Brot.) Jus
Page 275 and 276:
against this name and this was the
Page 277 and 278:
Gyptis crassipes (Hieron.) R. M. Ki
Page 279 and 280:
Hebeclinium DC., Prodr. 5: 136 (183
Page 281 and 282:
Fisher, T. R. (1957). Taxonomy of t
Page 283 and 284:
2800-4000 m. January-April. *Helogy
Page 285 and 286:
Note: Robinson (2006: 187) was of t
Page 287 and 288:
Arvet-Touvet, C. (1881). Spicilegiu
Page 289 and 290:
adical leaves present at flowering
Page 291 and 292:
Tarija: Cuesta de Sama, 3000 m, Mey
Page 293 and 294:
Hieracium microcephalum Sch.Bip., B
Page 295 and 296:
La Paz: Unduavi, 3200 m, Buchtien 4
Page 297 and 298:
References Cabrera, A. L. (1951). N
Page 299 and 300:
Cochabamba: Wood 11849 (K), Wood 17
Page 301 and 302:
Cabrera, A. L. (1963). Estudios sob
Page 303 and 304:
Argentina, Bolivia (Cochabamba, La
Page 305 and 306:
Argentina, Bolivia (Chuquisaca, Coc
Page 307 and 308:
Hypochaeris taraxacoides [sic!] (Me
Page 309 and 310:
Ichthyothere terminalis (Spreng.) M
Page 311 and 312:
Santa Cruz: Wood 12753 (K), Wood et
Page 313 and 314:
J Jaegeria Kunth in Humb., Bonpl. &
Page 315 and 316:
Jaumea mimuloides Hieron., Bot. Jah
Page 317 and 318:
Jungia (Martrasia) affinis Gardner,
Page 319 and 320:
?Jungia jelskii Hieron., Bot. Jahrb
Page 321 and 322:
*Eupatorium gynoxioides Rusby, Bull
Page 323 and 324:
Koanophyllon simillima (B. L. Rob.)
Page 325 and 326:
Bolivia (Cochabamba, La Paz, Tarija
Page 327 and 328:
Laggera lyrata (Kunth) Leins, Mitt.
Page 329 and 330:
Lepidaploa remotiflora (Rich.) H. R
Page 331 and 332:
are of the opinion that it should b
Page 333 and 334:
Eizaguirrea Remy in Gay, Flora de C
Page 335 and 336:
Liabum pallatangense Hieron., Bot.
Page 337 and 338:
Moquinia macrocephala Rusby, Descr.
Page 339 and 340:
Type: Eupatorium viscidum Hook. & A
Page 341 and 342:
Loricaria thujoides (Lam.) Sch.Bip.
Page 343 and 344:
Lucilia lundii Baker in Mart., Fl.
Page 345 and 346:
Anderberg, A. A. & S. E. Freire. (1
Page 347 and 348:
*Lycoseris boliviana Britton, Bull.
Page 349 and 350:
Melampodium L. sect. Alcina (Cav.)
Page 351 and 352:
Lipotriche R.Br., Observ. Campos :
Page 353 and 354:
distributed with large sets of Kerb
Page 355 and 356:
Mesanthophora rojasii (Cabrera0 H.
Page 357 and 358:
Holmes, W. C. & S. McDaniel (1989).
Page 359 and 360:
Milkania meridana V. M. Badillo, Bo
Page 361 and 362:
[Original publication not seen.] Ci
Page 363 and 364:
Wet evergreen forest, river banks,
Page 365 and 366:
*Mikania haenkeana DC., Prodr. 5: 1
Page 367 and 368:
Bolivia (?), Brazil, Venezuela. Ver
Page 369 and 370:
Mikania napensis S. F. Blake, J. Wa
Page 371 and 372:
Mikania lima Schltdl. ex Mart., Flo
Page 373 and 374:
(01399088). Note: The K and US isot
Page 375 and 376:
Mikania vitrea B. L. Rob., Contr. G
Page 377 and 378:
*Montanoa orbignyana Klatt, Abh. Na
Page 379 and 380:
Paramo Yungeno. 2950 m. April-May.
Page 381 and 382:
Munnozia pulchra (S. F. Blake) H. R
Page 383 and 384:
3. (2) Apices of outer phyllaries r
Page 385 and 386:
Mutisia cneorifolia Dec. ex Wedd.,
Page 387 and 388:
Bolivia (Chuquisaca, Cochabamba, La
Page 389 and 390:
N *Nardophyllum armatum (Wedd.) Rei
Page 391 and 392:
Grassland, cerrado, semideciduous f
Page 393 and 394:
Gynoyxs alternifolia Sch.Bip., Bull
Page 395 and 396:
for the type. The name provided by
Page 397 and 398:
*Nardophyllum armatum (Wedd.) Reich
Page 399 and 400:
Onoseris integrifolia Less., Linnae
Page 401 and 402:
*Ophryosporus angustifolius B. L. R
Page 403 and 404:
Bolivia (Cochabamba). Scrub. 2500 m
Page 405 and 406:
Orthopappus Gleason, Bull. New York
Page 407 and 408:
Bolivia (Pando). 455 m. June. 407
Page 409 and 410:
Pappobolus S. F. Blake, Hooker‘s
Page 411 and 412:
Leaves 2-6 mm long, margins revolut
Page 413 and 414:
References Cabrera, A. L. (1978). P
Page 415 and 416:
Robinson, H. (2006). Pectis. In: G.
Page 417 and 418:
4. (3) Leaves 9-11 cm long by 3-4 c
Page 419 and 420:
specifically. Since second stage le
Page 421 and 422:
c. 2700 m. October. Pentacalia subg
Page 423 and 424:
Perezia abbiattii Cabrera Darwinian
Page 425 and 426:
Perezia hunzikeri Cabrera, Bol. Soc
Page 427 and 428:
*Perezia pungens (Humb. & Bonpl.) L
Page 429 and 430:
Psilopogon Phil., Linnaea 33(1): 12
Page 431 and 432:
Carphobolus Schott subgen. Eucarpho
Page 433 and 434:
Bolivia (Santa Cruz), Brazil. Cerra
Page 435 and 436:
Plazia daphnoides Wedd., Chloris An
Page 437 and 438:
Pluchea dodoneifolia (Hook. & Arn.)
Page 439 and 440:
Argentina, Bolivia (Santa Cruz), Br
Page 441 and 442:
Polyachyrus poeppigii Less. ex Poep
Page 443 and 444:
Pontesia Vell., Fl. Flum. 8: t. 147
Page 445 and 446:
*Porophyllum ruderale (Jacq.) Cass.
Page 447 and 448:
March. Note: Eupatorium erythrolepi
Page 449 and 450:
CHILI: endrots des montagnes d‘Ac
Page 451 and 452:
Pseudognaphalium Kirp. subgen. Laph
Page 453 and 454:
Pseudognaphalium graveolens (Kunth)
Page 455 and 456:
Pseudonoseris discolor (Muschl.) H.
Page 457 and 458:
Pterocaulon hassleri Chodat, Bull.
Page 459 and 460:
Ptilurus D. Don, Trans. Linn. Soc.
Page 461 and 462:
por San Juan del Potrero y bajando
Page 463 and 464:
Rhysolepis emaciata H. Rob. & A. J.
Page 465 and 466:
S Sabazia Cass., Dict. Sci. Nat. 46
Page 467 and 468:
Ariza Espinar, L. (2000). Sanvitali
Page 469 and 470:
Note: Robinson (2006) took a very b
Page 471 and 472:
Schkuhria bonariensis Hook. & Arn.,
Page 473 and 474:
Senecio L. sect. Dendrophorbium Cua
Page 475 and 476:
Plants stemmed with various capitul
Page 477 and 478:
Capitula few, laxly cymose-corymbos
Page 479 and 480:
The following specimens were cited
Page 481 and 482:
Lomanthus bangii (Rusby) B. Nord. &
Page 483 and 484:
Senecio campanulatus Sch.Bip., Bull
Page 485 and 486:
Senecio culcitioides Sch.Bip., Bonp
Page 487 and 488:
*Senecio glacialis (Meyen & Walp.)
Page 489 and 490:
Senecio pulviniformis Hieron., Bot.
Page 491 and 492:
*Senecio psychrophilus Wedd., Chlor
Page 493 and 494:
Note: The following specimens were
Page 495 and 496:
var. macbridei (Cuatrec.) Cabrera,
Page 497 and 498:
Senecio spinosus DC., Prodr. 6: 420
Page 499 and 500:
Senecio trifurcifolius Hieron., Bot
Page 501 and 502:
Seris Willd., Mag. Neuesten Entdeck
Page 503 and 504:
Siegesbeckia melanolepis Sch.Bip.,
Page 505 and 506:
Robinson, H. (2006). Smallanthus. I
Page 507 and 508:
Sobreyra oblonga Ruiz & Pav., Syst.
Page 509 and 510:
Solidago odora Ait. [as ‗Soldiago
Page 511 and 512:
Sonchus umbellifer Thunb., Fl. Cap.
Page 513 and 514:
Sonchus australis Hort. ex G. Trevi
Page 515 and 516:
Sparganophorus vaillantii Crantz, I
Page 517 and 518:
Sphagneticola ulei O. Hoffm., Notiz
Page 519 and 520:
Spilanthes tenella Kunth in Humb.,
Page 521 and 522:
3000-4000 m. March. Robinson (1932)
Page 523 and 524:
*Stevia chamaedrys Griseb., Abh. K
Page 525 and 526:
Note: Robinson (1931: 8) noted Tate
Page 527 and 528:
Bolivia (Santa Cruz). Ancient clear
Page 529 and 530:
Bolivia (Cochabamba). 3000 m. March
Page 531 and 532:
Poorly- to well-branched shrubs; pl
Page 533 and 534:
Gamochaeta capitata Wedd., Chloris
Page 535 and 536:
Tripolium moelleri Phil., Anales Un
Page 537 and 538:
Balansa collection in K gives ‗L
Page 539 and 540:
?Bolivia (?), Chile, ?Ecuador. Cite
Page 541 and 542:
*Tagetes maxima Kuntze, Revis. Gen.
Page 543 and 544:
Cochabamba: Buchtien 2399 (US); Jim
Page 545 and 546:
eferring to the place of publicatio
Page 547 and 548:
Tilesia baccata (L.) Pruski, Novon
Page 549 and 550:
Zardini, E. M. (1974). Dos compuest
Page 551 and 552:
Slopes, dry hillsides, Puna, loose
Page 553 and 554:
Bartolia Adans., Fam. Pl. 2: 124 (1
Page 555 and 556:
Castra Vell., Fl. Flum. : 342 (1825
Page 557 and 558:
15066 but is not marked as a Humbol
Page 559 and 560:
Trixis divaricata (Kunth) Spreng. v
Page 561 and 562:
U Ucacou Adans., Fam. Pl. 2: 131 (1
Page 563 and 564:
References Blake, S. F. (1925). On
Page 565 and 566:
Ximenesia microptera DC., Prodr. 5:
Page 567 and 568:
Verbesina microptera (DC.) Herter,
Page 569 and 570:
Vernonia Schreb. sect. Tephrodes DC
Page 571 and 572:
Vernonia Schreb. sect. Vernonia sub
Page 573 and 574:
Vernonia acilepis Benth. in Oersted
Page 575 and 576:
Vernonia odoratissima Kunth in Humb
Page 577 and 578:
Vernonia chrysophylla Gardner, Lond
Page 579 and 580:
Cerrado, sandy soils. 490-660 m. Ma
Page 581 and 582:
Bolivia (Santa Cruz), Brazil, Parag
Page 583 and 584:
March-May. Note: The maps in Dematt
Page 585 and 586:
Vernonia neriifolia Gardner, London
Page 587 and 588:
‗Mexico, Central America, Andean
Page 589 and 590:
Vernonia remotiflora Rich. var. ari
Page 591 and 592:
*Vernonia rufopapposa Hieron. var.
Page 593 and 594:
Mexico, Central America, Colombia,
Page 595 and 596:
Vernonia subobscura Malme, Ark. Bot
Page 597 and 598:
References Blake, S. F. (1918). A r
Page 599 and 600:
*Viguiera glutinosa Rusby, Mem. Tor
Page 601 and 602:
Viguiera stenophylla (Hook. & Arn.)
Page 603 and 604:
W Wahlenbergia Schumach., Beskr. Gu
Page 605 and 606:
Werneria Kunth subgen. Anactis (Gay
Page 607 and 608:
?Werneria juniperina Hieron., Bot.
Page 609 and 610:
Bolivia (La Paz, Potosí), Peru. Ma
Page 611 and 612:
Wullfia havanensis DC., Prodr. 5: 5
Page 613 and 614:
concept of X. spinosum given in thi
Page 615 and 616:
Xanthium orientale L. forma lacinia
Page 617 and 618:
Xanthium spinosum L. var. synacanth
Page 619 and 620:
Xanthium acutum Greene, Pittonia 4(
Page 621 and 622:
Xanthium mongolicum Kitag., Rep. Fi
Page 623 and 624:
Xenophyllum ciliolatum (A. Gray) V.
Page 625 and 626:
5200 m. November. Xenophyllum staff
Page 627 and 628:
Y Z Zandera D. L. Schulz, Haussknec
Page 629 and 630:
Zinnia multiflora L., Sp. Pl. (ed.
Page 631 and 632:
References Anderberg, A. A., Salter
Page 633 and 634:
Cabrera, A. L. (1935). Notas sobre
Page 635 and 636:
Descole, H. R., O‘Donell, C. A. &
Page 637 and 638:
Heering, W. (post May 1915 or early
Page 639 and 640:
Koster, J. Th. (1947). Plants colle
Page 641 and 642:
Robinson, B. L. (1906a). Studies in
Page 643 and 644:
Soó, R. (1967). Species et combina
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Checklist of Bolivian Compositae - Royal Botanic Gardens, Kew

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