102 SPECIES PLANTARUM — FLORA OF THE WORLD (2002) 54. <strong>Luzula</strong> taurica (V.I.Krecz.) Novikov, Nov. Sist. Vyssh. Rast. 27: 20 (1990) <strong>Luzula</strong> campestris subsp. taurica V.I.Krecz., Žurn. Russk. Bot. Obshch. Akad. Nauk S.S.S.R. 12: 490 (1928); <strong>Luzula</strong> multiflora subsp. taurica (V.I.Krecz.) Novikov, in Fl. Evr. Casti SSSR 2: 83 (1976). T: Crimea, Demerdzhi Yaila, 25 May 1905, N.A.Busch; lec<strong>to</strong>: LE, fide J.Kirschner, Taxon 39: 113 (1990). <strong>Luzula</strong> multiflora subsp. caucasica V.I.Krecz., Žurn. Russk. Bot. Obshch. Akad. Nauk S.S.S.R. 12: 490 (1928). T: [Georgia, Ahalcihe] ad fines inter Transcaucasiam rossicam et Turciae districtum Achalzich, 7 Jul 1874, G.F.Radde; syn: LE; [Georgia] Cartalinia, prope Turam [?], A.J.Szovits 25; syn: LE. <strong>Luzula</strong> campestris var. debilis Velen., Fl. Bulg. 572 (1891). T: Bulgaria, Mt Kom above Berkovica, Jul 1889, J.Velenovský, syn: PRC, W. <strong>Luzula</strong> campestris var. atrofusca Maly, Glasn. Zemaljsk. Muz. Bosni Hercegovini 32: 141 (1920). T: Bosnia, Sarayevo, Mt Gola Jahorina, K.Maly; syn: SARA. Perennials, usually 15–25 cm tall, densely or rarely loosely caespi<strong>to</strong>se; rhizome short, without s<strong>to</strong>lons. Basal leaves usually up <strong>to</strong> 8–10 cm long, (2.5–) 3.0–4.0 (–5.0) mm wide; cauline leaves 1–2, usually 2.5–4.0 cm long, 2–3 mm wide; all leaves with obtuse tip and densely papillose-serrulate margins. Inflorescence composed of 1 (–3) subsessile and 2–6 (–7) pedunculate ovoid <strong>to</strong> hemispherical (5–) 7–9 (–11)-flowered clusters; peduncles straight, rarely slightly flexuose, ±smooth, without secondary branches; longest peduncles up <strong>to</strong> 2–4 cm long. Lower bract usually 1–3 cm long, shorter than inflorescence. Bracteoles membranous, 1.5–2.5 mm long, ovate, fimbriate-ciliate above. Tepals ±equal, entire, acuminate, usually 2.9–3.5 mm long, dark <strong>to</strong> blackish brown, with indistinct paler brownish margins. Stamens 6; anthers (0.8–) 1.1–1.3 (–1.4) mm long; filaments c. 0.5–0.6 mm long; style 0.7–1.1 mm long; stigmas 1.8–3.0 mm long, subpersistent. Capsule ±obovoid, ±shorter than perianth, pale brown <strong>to</strong> dark brown; capsule segments 2.4–3.0 mm long. Seeds 1.0–1.2 mm long, 0.7–0.8 mm wide; appendage (0.2–) 0.3–0.4 (–0.5) mm long. 2n=12 [12AL], fide J.Kirschner, Thaiszia 2: 30–31 (1992). Fig. 22. Balkan Peninsula and Crimea in S Europe, Turkey, and Georgia and Armenia in the Caucasus. 13: ALB, BUL, GRC, ROM, TUE, YUC. 14: KRY. 33: NCS, TCS. 34: TUR. Subalpine meadows and grasslands. Map 85. 13: ALBANIA, Skutari, Zukali, 15 Jul 1897, A.Baldacci [It. Alban. 5] 47 (K); Vermoshe,‘Gipfelregion von Grebeni Selces’, 1700 m, I.Dörfler 283 (G, K, M, WU). BULGARIA: Rila Mtns, Rilski Monastyr, Dolgia Rid, J.Štěpánková s.n. (PRA). GREECE: Serrai, Mt Vrondous, 1750–1849 m, S.Snogerup & A.Strid 5004 (LD, PRA). ROMANIA: Hunedoara, Retezat, Mt Borescu, 12 Jul 1924, E.J.Nyárády et al. [Fl. Rom. Exs.] 933 (K). F.Y.R. MACEDONIA: Vodno, 2 May 1972, R.Bělohlávková (PRA); Shar Planina, Kobilica, 2520 m, 27 Aug 1938, B.Pawlowski (KRAM). 14: CRIMEA, Yalta, Nikitsky Yaila, 1350 m, P.H.Davis 33387 (K); Romak-Kosh, Babugan-Yaila, 20 Jul 1948, S.V.Yuzeptshuk (LE); Alushta, Luchis<strong>to</strong>e, Mt Demerdzhi, 1100–1200 m, J.Kirschner L189 (PRA). 34: TURKEY: Trabzon, Karakö, N side of Pontus Pass, 2100 m, E.Hennipman et al. 1718 (K). Similar <strong>to</strong> L. abchasica and L. campestris. From the latter it differs in caespi<strong>to</strong>se growth and seed characters. A very variable species (fide J.Kirschner, 1993: 170–172); the isolated populations differ from one another in a number of minor quantitative traits. 55. <strong>Luzula</strong> divulgata Kirschner, Folia Geobot. Phy<strong>to</strong>tax. 14 [1979]: 431 (1980) T: Czech Republic, Moravia, between Sobůlky and Věteřov, 6 May 1978, J.Kirschner s.n.; holo: PRC539. Illustration: J.Kirschner, Folia Geobot. Phy<strong>to</strong>tax. 28: 159, fig. 6 (1993). Perennials, (10–) 15–25 (35) cm tall, densely caespi<strong>to</strong>se, hairy; s<strong>to</strong>lons absent; rhizome short. Basal leaves usually 10–15 cm long, 3.0–5.5 mm wide; cauline leaves usually 2 or 3, usually 3.5–<strong>6.</strong>5 cm long, 3–5 mm wide; all leaves with obtuse tip and sparsely papilloseserrate margin. Lower bract usually 1.6–3.0 cm long, shorter than inflorescence. Inflorescence composed of one subsessile and 2–8 pedunculate clusters; clusters broadly ovoid <strong>to</strong> hemispherical, 6–11-flowered. Peduncles straight or flexuose, smooth, without secondary branches. Longest peduncles (including clusters) 2.1–5.0 cm long. Bracteoles 1.8–2.4 mm long, fimbriate-ciliate above. Tepals ±equal, entire, ±acuminate, usually 3.5–4.7 mm long, castaneous brown <strong>to</strong> dark brown. Stamens 6; anthers usually 2.3–3.3 mm long; filaments 0.5–0.7 mm long; ovary 1.0–1.7 mm long; style usually 1.7–1.9 mm long; stigmas subpersistent, 2.5–4.5 mm long, conspicuously exserted. Capsule brown, oblong-ovoid, not
<strong>JUNCACEAE</strong> (<strong>Luzula</strong>) Figure 22. <strong>Luzula</strong> multiflora subsp. multiflora. A mountain form. A, habit; B, flower cluster. <strong>Luzula</strong> campestris subsp. campestris. C, habit; D, seeds; E, perianth. <strong>Luzula</strong> taurica. F, habit. (A, B, Z.Skála & H.Skálová L338, PRA; C–E, J.Kirscher L501, PRA; F, J.Kirschner L189, PRA). Scale bars: A, C, F = 5 cm; B = 5 mm; D = 1 mm; E = 2 mm. Drawn by © E.Smrčinová. 103
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