Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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58 SPECIES PLANTARUM — FLORA OF THE WORLD (2002) 7: Tepals ±glabrous (not to be confused with ciliate bracteoles) 9 Inflorescence usually distinctly nodding 10 Style 0.1–0.2 mm long; capsule segments 1.5–1.8 mm long; leaf margin ±densely papillose-serrulate 40. L. racemosa 10: Style 0.3–0.4 mm long; capsule segments 1.8–2.1 mm long; leaf margin ±smooth 26. L. traversii 9: Inflorescence erect to suberect 11 Inflorescence composed of 3–6 (10) distinct elongated cylindrical flower clusters; lower clusters pedunculate and upper subsessile 37. L. brachyphylla 11: Inflorescence congested, usually a single head, ovoid to lobate, rarely interrupted at base; clusters ±sessile 12 Basal leaves 4–9 mm wide; basal bract herbaceous; style c. 0.3 mm long. [Juan Fernández Is.] 35. L. masafuerana 12: Basal leaves c. 1 mm wide; basal bract membranous; style c. 0.1 mm long. [NW Argentina] 44. L. ruiz-lealii 4: Stamens 6, rarely 4 or 5 13 Tepals densely to sparsely ciliate 36. L. alopecurus 13: Tepals ±glabrous 14 Seeds (excluding basal appendage) more than 1.2 mm long 15 Seeds 2.1–2.4 mm long 30. L. pediformis 15: Seeds less than 2.0 mm long 16 Anthers 2.0–2.9 mm long, at least 5 times as long as filaments 31. L. caespitosa 16: Anthers less than 1.2 (–1.5) mm, up to 3 times as long as filaments 17 Style 0.6–0.9 mm. [S Europe] 28. L. pindica 17: Style 0.3–0.6 mm. [South America] 18 Tepals ±equal, c. 2.7–3.3 mm long; anthers usually 0.6–0.9 mm long; inflorescence ±compact 38. L. chilensis 18: Tepals unequal, with outer ones 3.5–4.8 mm long; anthers usually 0.7–1.5 mm long; inflorescence often interrupted at base 39. L. leiboldii 14: Seeds (excluding basal appendage) less than 1.2 mm long 19 Inflorescence erect to suberect 20 Inflorescence of a single compact ±ovoid head; cauline leaves absent; basal leaves ±obtuse; basal bract membranous, to 1 cm long 44. L. ruiz-lealii 20: Inflorescence composed of 3–6 (–10) distinct spike-like elongated cylindrical flower clusters; lower clusters pedunculate and upper ones subsessile; cauline leaves 1 or 2; basal bract herbaceous, 1.5–4.0 cm long 37. L. brachyphylla 19: Inflorescence nodding 21 Inflorescence a panicle of remotely interrupted partial inflorescences, each of long pedunculate secondary panicles composed of 3–10 spike-like clusters; lowermost secondary panicle often 10–15 cm below the terminal one 34. L. mendocina 21: Inflorescence congested in a single head, sometimes lobate or interrupted at base; basal clusters subsessile, not remote
JUNCACEAE (Luzula) 22 Leaf margin ±smooth; basal leaves acute with acicular tip 22: Leaf margin ±densely papillose-serrulate; basal leaves ±obtuse 26. L. traversii to acute (on a single plant) 29. L. spicata 25. Luzula ulophylla (Buchenau) Cockayne & Laing, Trans. Proc. New Zealand Inst. 43: 366 (1911) Luzula racemosa var. ulophylla Buchenau, Oesterr. Bot. Z. 48: 245 (1898). T: New Zealand, South Island, Albertown, Lake Wanaka, D.Petrie; syn: W [seen in 1994, in 1998 not located]; Southern Canterbury Alps, Castle Hill Distr., 2200 ft., Dec 1894, L.C.Cockayne; syn: B, destroyed; Castle Hill, 1893, L.C.Cockayne; neo: WELT 12668, fide E.Edgar, New Zealand J. Bot. 4: 165 (1966); E.Edgar, in L.B.Moore & E.Edgar, Fl. New Zealand 2: 72 (1970). Perennials (5–) 9–20 (–28) cm long, densely hairy, densely caespitose; rhizome short, vertical, ±many-headed; stolons absent. Basal leaves short, usually 3–6 cm long, (1–) 2–3 mm wide, usually inrolled, narrow, whole, or upper two thirds of abaxial surface with thick white tomentum; cauline leaves absent or 1 (–2), with blade reduced, filiform, 1–2 cm long; sheath to 2 cm long. All leaves with subulate acute tip; margins ±smooth. Lower bract ovate, strawbrown, membranous, c. 5 mm long, similar to middle bracts. Inflorescence erect, cylindrical to ±ovoid, densely congested, spike-like, usually 0.8–2.0 × 0.6–1.0 cm, of 1–5 few-flowered clusters. Bracteoles ovate, scarious, 1.5–2.0 mm long, ciliate near apex. Tepals ±entire, subequal, rarely ±denticulate near apex, (1.3–) 1.6–1.9 (–2.1) mm long, acuminate; lower middle part dark castaneous brown; margins broad, white-scarious. Stamens 3 (4–6); anthers 0.4–0.5 (–0.7) mm long; filaments 0.3–0.5 mm long; style c. 0.3 mm long; stigmas c. 0.5 mm long. Capsule obovoid, equalling or slightly exceeding perianth, ±obtuse to subacute, dark brown to black; capsule segments 1.5–2.0 × c. 1.1–1.3 mm. Seeds pale castaneous, 0.8–0.9 mm long, c. 0.6 mm wide; appendage minute, appearing as a paler basal fibrillate swelling to 0.1 mm long. 2n=48 [48CL], fide H.Nordenskiöld, New Zealand J. Bot. 4: 186 (1966); 2n=44, fide J.B.Hair, New Zealand J. Bot. 5: 18 (1967). Fig. 13. Endemic to the South Is. of New Zealand. 51: NZS. Wind-eroded shingly open ground or grasslands, between 300 and 1500 m. Map 50. 51. NEW ZEALAND SOUTH: South Is., Otago, head of Eweburn Dam, H.E.Connor & H.Nordenskiöld S151 (UPS); C Otago, Queenstown, Ben Lomond, H.E.Connor & H.Nordenskiöld S189 (UPS); Canterbury, Acheron R., near diversion to Lake Coleridge, E.Edgar G5791 (CHR, PR); S Canterbury, Lake Tekapo, Boundary Stream, 13 Jan 1961, H.E.Connor (CHR 113675, K); Canterbury, Lake Heron, 720 m, A.D.Wilton 98048 (CHR, PRA). Easily distinguished by the dense white tomentum on the abaxial surface of the leaves. 26. Luzula traversii (Buchenau) Cheeseman, Man. New Zealand Fl., 2nd edn, 305 (1925) Luzula racemosa var. traversii Buchenau, Bot. Jahrb. Syst. 12: 133 (1890). T: New Zealand, H.H.Travers; lecto: WELT 12718, fide E.Edgar, New Zealand J. Bot. 4: 164 (1966); E.Edgar, in L.B.Moore & E.Edgar, Fl. New Zealand 2: 71 (1970). Perennials 10–40 cm tall, densely caespitose; rhizome short; stolons absent. Leaves with ±acicular tip and sparsely ciliate, with ±smooth margins. Lower bract herbaceous, exceeding inflorescence, 3–5 cm long. Inflorescence ±nodding, of (1–) 3–9 clusters congested into a lobate or pyramidal head up to 2–3 × 1.5 cm. Bracteoles ovate, entirely membranous, often castaneous at base, 1.5–2.0 mm long, ±entire or sparsely ciliate near apex. Tepals subequal, ±entire, usually with a dark castaneous brown midrib and broad whitish-membranous margins. Style 0.3–0.4 mm long; stigmas c. 1.0–1.2 mm long. Capsule subacute, equalling or ±exceeding perianth. Seeds 1.0–1.2 mm long, 0.6–0.7 mm wide; appendage indistinct, to 0.1 mm, fibrillate. Endemic to the South Is. of New Zealand. Similar to L. racemosa but inflorescence not interrupted, tepals with broad scarious-membranous margins, and longer style, capsule segments and seeds. L. masafuerana differs from L. traversii in having broader leaves with papillose-serrulate margins, broader bracts and suberect inflorescence. Two varieties are recognized within L. traversii; intermediates are known (Canterbury, Cave Stream, N.T.Moar & H.Nordenskiöld S118, UPS). E.Edgar, New Zealand J. Bot. 4: 164 (1966), mentioned 59
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AUSTRALIAN BIOLOGICAL RESOURCES STU
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CONTENTS Contributors iv Species Pl
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Species Plantarum Steering Committe
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PREFACE This volume is the first of
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