Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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82 SPECIES PLANTARUM — FLORA OF THE WORLD (2002) 3–4 cm long, c. 1 mm wide, with ±obtuse to subobtuse tip and margins ±smooth; cauline leaves absent. Basal bract pale straw-brownish membranous, oblong-lanceolate, acute, ciliate, up to 1 cm long. Inflorescence spike-like, densely congested, terminal, cylindricalovoid,1.0–1.5 × c. 0.7–0.8 cm, of 2–6 few-flowered clusters, or rarely of 1–3 remote basal clusters. Bracteoles whitish scarious, ovate, acuminate, ciliate, to 1.5–1.8 mm long. Tepals glabrous, ±equal, pale straw-brown or membranous above, brown below, acuminate (subaristate), with inner tepal ±mucronate, usually 2.1–2.6 mm long. Stamens 3–5 (–6); anthers 0.4–0.6 mm long; filaments 0.4–0.6 mm long; style c. 0.1 mm long; stigmas to 1 mm long. Capsule ovoid, subacute, paler brown, shorter than perianth; capsule segments 1.6–1.8 × c. 1.0 mm. Seeds 1.0–1.1 mm long, c. 0.6 mm wide; appendage yellowish, c. 0.2 mm long, fibrillate. Endemic to Mendoza, Argentina. 85: AGW-ME. Open patches in dry grasslands, above 3000 m. Map 74. 85. ARGENTINA NORTHWEST: Mendoza, San Carlos, Laguna del Diamante, 3200 m, 16 Jan 1952, A.Ruíz Leal 14584 (SI); 16 Jan 1949, A.Ruíz Leal 11717 (LIL264113, MERL); Mendoza, Tunuyán, Valle del Alto Tunuyán, Cerro Pan de Azúcar, 6 Feb 1934, A.Ruíz Leal 2140 (LIL, MERL). A peculiar dwarf form, superficially similar to L. ulophylla of New Zealand. Number of stamens varies within inflorescence and cannot be used to separate L. ruiz-lealii and L. parvula. Suspected to be a depauperate form parallel to that of L. racemosa (L. macusaniensis) but cannot be equated with any of the southern hexandrous forms. Doubtful names: Luzula psilophylla Phil., Linnaea 33: 267 (1865) Luzula chilensis f. psilophylla (Phil.) Fuentes, Bol. Mus. Nac. Hist. Nat. 90: 138 (1917). T: In Andibus ‘Talcaregue’. [J.Gay]; holo: n.v. The type material has not been located. The protologue description does not give relevant information, and later herbarium material annotated by Philippi is taxonomically heterogenous, including Luzula leiboldii, L. chilensis and L. brachyphylla. Luzula chilensis var. pallida Buchenau ex Fuentes, Bol. Mus. Nac. Hist. Nat. 90: 138 (1917) T: [not indicated]. Presumed Hybrids Luzula ×somedana Fern.-Carv. & Fern.Prieto in J.A.Fernández Prieto & M. de C.Fernández- Carvajal, Stud. Bot. 2: 134 (1983) = Luzula pediformis (Chaix) DC. × L. sylvatica subsp. henriquesii (Degen) Pirajá T: Spain, Asturias, Somiedo, Sierra del Parámo, Laguna Cavera, 1720 m, 21 Jul 1978, J.A.Fernández-Prieto; holo: FCO8195; see also J.A.Fernández Prieto & C.Fernández-Carvajal, Stud. Bot. 2: 136, fig. A, B, C (1983). A seed-sterile hybrid. Luzula ×gayana Font Quer & Rothm., Cavanillesia 7: 175 (1936) = Luzula pediformis (Chaix) DC. × L. caespitosa (E.Mey.) Steud. T: Spain, Asturias, Cueto de Arvas, 1700 m, 15 Jul 1935, P.Font Quer & W.Rothmaler; holo, n.v. Material not seen but the existence of the hybrid is probable. Luzula ×marceloi Rivas Mart., Anales Inst. Bot. Cavanilles 21(1): 289 (1963), nom. inval. = Luzula caespitosa (E.Mey.) Steud. × L. spicata (L.) DC. T: Spain, Sierras de Guadarrama y Gredos, Ladera del Noruego, 2200 m, S.Rivas Martínez; holo: n.v.
JUNCACEAE (Luzula) Sect. 6. THYRSANOCHLAMYDEAE Luzula sect. Thyrsanochlamydeae Satake, in T.Nakai & M.Honda, Nov. Fl. Jap. 1: 30 (1938) T: Luzula unalaschkensis (Buchenau) Satake; lecto, designated here. Luzula subsect. Thyrsanochlamydeae Novikov, Byull. Moskovsk. Obshch. Isp. Prir., Otd. Biol., 95(6): 67 (1990), nom. illeg. T: Luzula confusa Lindeb. Luzula subsect. Tundricolae Kovt., Bot. Žurn. (Moscow & Leningrad) 72: 1399 (1987). T: Luzula tundricola Gorodkov ex V.N.Vassil. Luzula subsect. Confusae Kovt., Bot. Žurn. (Moscow & Leningrad) 72: 1399 (1987). T: Luzula confusa Lindeb. Luzula subsect. Nivales [Chrtek & Křísa ex] Kovt., Bot. Žurn. (Moscow & Leningrad) 72: 1399 (1987). T: Luzula nivalis (Laest.) Spreng. Perennials. Leaves with tip acute, acuminate or mucronate. Lower bract short, membranous to brownish-membranous. Inflorescence subcongested, or in glomerules on thin peduncles which are usually arcuate or nodding, less often straight. Seeds ellipsoidal to narrowly ellipsoidal; appendage basal, short, up to 0.2 mm long, or lacking; base fibrillate. Five species, almost confined to the Arctic and rare in the mountains in northern temperate regions. They represent a link between sect. Diprophyllatae and the type section. In particular, L. arcuata and L. confusa may have an alloploid origin of the above parentage, as indicated by their karyotypes (Nordenskiöld, 1949, 1951). 1 Basal leaves usually 4.5–6.0 mm wide, bluish green; plants usually tall, 20–40 cm tall [California] 45. L. subcongesta 1: Basal leaves usually narrower than 4 mm; plants usually less than 20 cm tall [circumpolar but north of California] 2 Basal leaves with obtuse, often slightly swollen tip with a spinuliform mucro; seed base with appendage c. 0.2 mm long and fibrillate 49. L. nivalis 2: Basal leaves subabruptly to gradually narrowing to acute or acuminate tip; seed base with short (to 0.1 mm long) appendage or without appendage, and always fibrillate 3 Plant base straw-brown to dull brownish; flower clusters pale brown to straw-brown, ellipsoidal 46. L. kjellmaniana 3: Plant base brown to purplish or reddish, usually shining; flower glomerules or clusters dark to blackish brown, ±globular or ellipsoidal 4 Peduncles c. 0.2 mm in diam., with secondary branches, usually arcuate to nodding; glomerules (1–) 2–6 (–8)-flowered, broader than wide; number of glomerules usually 5–10; stem less than 0.5 mm in diam. below inflorescence 47. L. arcuata 4: Peduncles c. 0.3 mm in diam., without secondary branches, usually straight, erect, less often arcuate; subsessile clusters more than 10-flowered; number of clusters usually 1–3; stem 0.5–0.9 mm in diam. below inflorescence 48. L. confusa 45. Luzula subcongesta (S.Watson) Jeps., Fl. Calif. 1: 258 (1921) Luzula spadicea var. subcongesta S.Watson, Bot. California 2: 202 (1880); Luzula parviflora var. subcongesta (S.Watson) Buchenau, Bot. Jahrb. Syst. 12: 110 (1890); Juncoides parviflora var. subcongesta (S.Watson) E.Sheld., Bull. Geol. Nat. Hist. Surv. 9: 63 (1894); Juncoides subcongesta Coville, Muhlenbergia 1: 105 (1904). T: California, Sierra Nevada, near Donner Lake, J.Torrey; syn: n.v. Illustration: Jepson, Fl. Calif. 1: 259, fig. 45c, d, e (1921). Map: J.Coffey Swab, Fl. North Amer. 22: 267 (2000). Perennials, loosely tufted, subglabrous, bluish green, with horizontal rhizome. Stem 20–40 cm long; cauline leaves 3–4, usually 3–5 cm long, 3–6 mm wide; basal leaves up to 10 (–15) cm 83
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AUSTRALIAN BIOLOGICAL RESOURCES STU
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CONTENTS Contributors iv Species Pl
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Species Plantarum Steering Committe
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PREFACE This volume is the first of
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2 SPECIES PLANTARUM — FLORA OF TH
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