Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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48 SPECIES PLANTARUM — FLORA OF THE WORLD (2002) appendage indistinct; base fibrillate. n=12 (2n=24); fide J.Pojar, Canad. J. Bot. 51: 625 (1973). Fig. 11. Endemic to SW Canada (Alberta and British Columbia) and Pacific NW of USA, from Washington and Montana to Oregon. 71: ABT, BRC. 73: IDA, MNT, ORE, WAS. Montane coniferous woodlands near timberline to subalpine and alpine slopes, ridges, meadows. Map 39. 71. BRITISH COLUMBIA: Manning Park, J.A.Calder & D.B.O.Savile 11642 (DAO, WTU). 73. MONTANA: Glacier National Park, B.Maguire & Piranian 15397 (WTU, UC). OREGON: Crater Lake National Park, Baker 6124 (UC). WASHINGTON: Mt Paddo, 5 Sep 1904, W.Suksdorf 4309 (L). Not located: 'from Fort Colville to Rocky Mts., 49° N', 1861, D.Lyall (P). 20. Luzula glabrata (Hoppe) Desv., J. Bot. (Desvaux) 1: 143 (1808) Juncus glabratus Hoppe, Herb. Viv. Pl. Rar. Praes. Alp., cent. 3, sine no. (1800); Juncus intermedius Host, Icon. Descr. Gram. Austriac. 3: 65 (1805), nom. illeg.; Luzula intermedia [Host] Baumg., Enum. Stirp. Transsilv. 3: 330 (1816), nom. illeg., non Hornem. (1815); Luzula spadicea var. glabrata (Hoppe) E.Mey., Syn. Luzul. 8 (1823); Luzula parviflora var. intermedia [Host] Wahlenb., Fl. Suec. 217 (1824), nom. illeg.; Luzula glabrata var. vera Buchenau, Krit. Verz. Juncac. 83 (1879), nom. inval.; Juncoides glabrata (Hoppe) E.Sheld., Bull. Geol. Nat. Hist. Surv. 9: 63 (1894). T: Austria, Salzburg, Untersberg, D.H.Hoppe; lecto: MW, fide L.Hämet-Ahti, Ann. Bot. Fenn. 8: 371 (1971); isolecto: PR. Juncus pilosus [Abart] C. glabrescens Schrank, Baier. Fl. 1: 621 (1789); Luzula glabrescens (Schrank) Dalla Torre & Sarnth., Farn- u. Blütenpfl. Tirol. 1: 434 (1906), nom. superfl. T: [Austria] 'aus dem Salzburgischen [Alpen]', K.M.Moll; syn: n.v. Illustrations: N.T.Host, Icon. Descr. Gram. Austriac. 3: plate 99 (1805); S.Pignatti, Fl. Ital. 3: 445 (1982). Perennials, usually 15–30 cm tall, loosely caespitose, subglabrous to ±glabrous; rhizome creeping to ascending; stolons usually ascending. Leaves of sterile shoots usually 10–15 cm long, (4–) 5–7 (–10) mm wide; leaves at stem base usually short, 2–5 cm long; cauline leaves 3–4, usually 3.5–6.0 cm long, 4.5–8.0 mm wide. All leaves with papillose-serrulate margins; apex acute. Lower bract ±herbaceous, to 2–3 cm long, much shorter than inflorescence. Inflorescence ±loose, ±nodding to suberect, ±divaricate, usually 5–9 × 3–6 cm, usually more than 35–50-flowered; flowers usually borne singly. Bracteoles ±ovate, membranous, often suffused brownish, c. 1.5–2.0 mm long, ciliate. Tepals equal, glabrous, ±lanceolate, acuminate, 2.4–3.0 mm long, deep to dark castaneous-brown, with margins indistinct or narrow, paler brownish. Stamens 6; anthers (1.1–) 1.4–1.6 (–2.1) mm long, up to 10 times as long as filaments; filaments 0.2–0.3 mm long; style 0.5–0.9 mm long; stigmas 1.0–1.5 mm long. Capsule ±ovoid, acute, shortly rostrate, dark brown, usually slightly exceeding perianth; capsule segments 2.5–2.9 × c. 1.4–1.6 mm. Seeds ±oblong-ovoid, c. 1.2–1.4 mm long, c. 0.7 mm wide; appendage indistinct; base fibrillate. 2n=12 [AL], fide H.Nordenskiöld, Hereditas 37: 330–331 (1951). Confined to the E Alps, from Berchtesgadener Alpen, Germany, to N Italy. 11: AUT, GER. 13: ITA. Alpine grasslands, stony slopes, usually above (1700–) 2000 m. Map 40. 11. AUSTRIA: Niederösterreich, Raxalpe, 26 Jul 1921, G.E. Du Rietz (UPS); Niederösterreich, Schneeberg, 1800–2050 m, G.Beck [Fl. Exs. Austro-Hung.] 1871/II (H, PR, SAM, W, WU); N Salzburg, Lofer, A.Spitzel [H.G.Reichenbach, Fl. Germ. Exs.] 1126 (H, K, PR). GERMANY: Bavaria, Berchtesgaden, Hohe Göll, Aug 1895, T.Wulff (LD); Berchtesgadener Alpen, Aug 1853, A.Progel (PR). 21. Luzula desvauxii Kunth, Enum. Pl. 3: 304 (1841) Luzula spadicea var. desvauxii (Kunth) E.Mey., Linnaea 22: 400 (1849); Luzula glabrata var. desvauxii (Kunth) Buchenau, Krit. Verz. Juncac. 83 (1879); Luzula glabrata subsp. desvauxii (Kunth) K.Richt., Pl. Eur. 1: 185 (1890); Luzula glabrata proles desvauxii (Kunth) P.Fourn., Quatre Fl. France 153 (1935). T: France, Auvergne, Mont Dore, Maire; holo: B, destroyed. Perennials usually (25–) 35–40 (–65) cm tall, loosely caespitose, subglabrous; rhizome creeping, branched; stolons short, ascending. Leaves of sterile shoots usually 15–20 cm long, 3–6 mm wide; basal leaves few, intermediate between cataphylls and cauline ones; cauline leaves 3–5, usually 10–20 cm long, 3.0–5.5 (–7.5) mm wide, conspicuously overlapping, with upper ones sometimes exceeding inflorescence. All leaves almost glabrous, with
JUNCACEAE (Luzula) papillose-serrulate margins; apex acute. Lower bract herbaceous, usually 2.5–5 cm long, shorter than or almost equalling inflorescence. Inflorescence many-flowered, ±nodding, later suberect, usually 5–7 × 4–5 cm; flowers borne singly or crowded in 2 (3–4)-flowered groups. Bracteoles scarious, ovate, ciliate, c. 1.5 mm long. Tepals equal, glabrous, lanceolate, acuminate, usually 2.4–2.8 mm long, deep to dark castaneous-brown; margins inconspicuous, paler (straw-) brown. Stamens 6; anthers (1.0–) 1.1–1.4 (–1.7) mm long; filaments 0.3–0.5 mm long; style c. 0.8–1.0 mm long; stigmas 1.0–1.5 mm long. Capsule oblong-ovoid, ±acute, shortly mucronate, dark brown, shining, ±equalling, or slightly longer than perianth; capsule segments (2.4–) 2.5–2.8 (–3.0) × c. 1.3–1.5 mm. Seeds ±ovoid, 1.1–1.3 mm long, c. 0.8 mm wide, brown; appendage to 0.1 mm long, fibrillate. 2n=12 [AL], fide A.-M.Lambert & J.Giesi, Inform. Annuelles Caryosyst. Cytogén. 1: 28 (1967). SW Europe, disjunctly in the E Pyrenees, Massif Central, Vosges and S Schwarzwald. Reported from Montes Cantábricos, N Spain, but material not seen [see P.Montserrat, Anales Inst. Bot. Cavanilles 21(2): 459-461 (1964)]. 11: GER. 12: FRA, SPA. Subalpine stony meadows, along streams, usually on non-calcareous substrates above 1200 m. Map 41. 11. GERMANY: Schwarzwald, Belchen, Jul 1858, H.A. de Bary (W). 12. FRANCE: Vosges, Hohneck, I.S.Perrin [F.W.Schultz, Herb. Norm.] 1154 (PR); Puy-de-Dôme, Mont-Dore, Grande Cascade, 1200–1300 m, 17 & 22 Jun 1971, Rastetter (H, LD); Auvergne, Puy-Mary, 1200 m, 28 Jul 1909, J.J.Héribaud (PR, W); Auvergne, Puy-Ferrand, 2 Sep 1852, E.Desvaux (H); Hautes Cévennes, S of Mt Mézenc, 1400 m, N.Y.Sandwith 3897 (K); E Pyrenees, Vallée d'Eyne, c. 2100 m, E.Wängsjö & G.Wängsjö 3366 (LD); E.M.Sennen [Pl. Esp.] 4055 (LD); E Pyrenees, L'Hospitalet, Ellman & N.Y.Sandwith 247 (K). SPAIN: E Pyrenees, Nuria, 1850 m, E.M.Sennen [Pl. Esp.] 1809 (BC, W). 22. Luzula parviflora (Ehrh.) Desv., J. Bot. (Desvaux) 1: 144 (1808) Juncus parviflorus Ehrh., Beitr. Naturk. 6: 139 (1791); Luzula spadicea var. parviflora (Ehrh.) E.Mey., Linnaea 22: 402 (1849); Juncoides spadicea var. parviflora (Ehrh.) Kuntze, Revis. Gen. Pl. 2: 724 (1891); Juncoides parviflora (Ehrh.) Coville, Contr. U.S. Natl. Herb. 4: 209 (1893). T: [Sweden, Lappland], C.Linnaeus; lecto: Flora Lapponica 124, Institut de France, Paris, designated here; photo BM, PRA. Luzula parviflora var. densiflora Lange, Consp. Fl. Groenland. 125 (1880). T: [not indicated; numerous syntypes from E and W Greenland]. Luzula parviflora var. sparsiflora Lange, Consp. Fl. Groenland. 125 (1880). T: [not indicated; numerous syntypes from Greenland]. Map: E.Hultén & M.Fries, Atlas N. Eur. Vasc. Pl. 1: map 166 (1986). Perennials 30–90 cm tall, loosely caespitose; rhizome ascending; stolons short, ascending, less often absent. Basal leaves 10–20 cm long, usually 5–10 mm wide; cauline leaves (3–) 4– 5, 3–10 mm wide. All leaves with papillose-serrulate margins; apex acute. Lower bract herbaceous or brownish membranous, usually 1.5–4.0 cm long, shorter than inflorescence. Inflorescence usually lax, sometimes nodding, with some main branches to 13 cm long, often arcuate; flowers borne singly. Bracteoles subglabrous to sparsely ciliate, ovate, membranous. Tepals equal, glabrous, ±lanceolate, 1.8–2.3 (–2.5) mm long. Stamens 6; anthers 0.3–0.6 mm long, as long as or shorter than filaments. Capsule smooth above. Seeds ±ellipsoid, brown to dark brown, 1.0–1.4 mm long; base fibrillate. A widely distributed northern circumpolar species, most common in subarctic areas and in the mountains of N Europe, N Asia and temperate North America. Three subspecies are recognized. 1 Inflorescence many-flowered with long main branches and short most distal secondary branches (peduncles); perianth segments medium brown 22a. subsp. parviflora 1: Inflorescence few-flowered, with long main branches and long distal secondary branches (almost divaricate); perianth segments usually pale brown to stramineous 2 Stem conspicuously reddish at base; cauline leaves mid-green, shining, tepals with a distinct midrib 22b. subsp. fastigiata 2: Stem without reddish coloration or with indistinct reddish tinge at base; cauline leaves bluish or greyish green, dull, tepals with an indistinct midrib 22c. subsp. melanocarpa 49
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AUSTRALIAN BIOLOGICAL RESOURCES STU
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CONTENTS Contributors iv Species Pl
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