Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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SPECIES PLANTARUM — FLORA OF THE WORLD (2002)
51. NEW ZEALAND SOUTH: South Is., Marlborough, Upper Awatere Valley, Muller Station, L.B.Moore
(CHR77227); Ashburton R., Blowing Point, A.Wall (CANTY1737); Dunedin, Hoopers Inlet, L.B.Moore
(CHR120151); Otago, Cromwell, Chafer Beetle Reserve, 210 m, A.D.Wilton 98223 (CHR, PRA); Otago,
Crystall's Beach, Cooks Head, A.D.Wilton 98019 (CHR, PRA); Stewart Is. Mason Bay, E of Big Sandhill,
H.Wilson (CHR).
Known from about 15 sites in the South Island and one site in Stewart Is.; vulnerable. This
species is extremely elusive, and is easily overlooked.
28. Luzula pindica (Hausskn.) Chrtek & Křísa, Webbia 19: 6 (1964); Notes Roy. Bot.
Gard. Edinburgh 25: 164 (1964)
Luzula spicata β [unranked] pindica Hausskn., Mitt. Thür. Bot. Ver. 13–14: 33 (1899); Luzula bulgarica
subsp. pindica (Hausskn.) Chrtek & Křísa, Bot. Not. 115: 308 (1962); Luzula spicata subsp. pindica
(Hausskn.) Gamisans, Candollea 29: 42 (1974). T: Greece, Pindhos Mtns, Mt Zygos, Jul 1885,
H.C.Haussknecht; syn: JE, W.
Illustration: J.Chrtek & B.Křísa, Bot. Not. 115: 309, fig. 7 (1962).
Maps: J.Chrtek & B.Křísa, Bot. Not. 115: 307, fig. 6 (1962); J.Chrtek & B.Křísa, Webbia 19: 8 (1964).
Perennials (15–) 20–35 (–45) cm tall, densely caespitose, sparsely hairy; rhizome short,
vertical; stolons absent. Basal leaves ±flat, c. 3.5–6 (–9) cm long, of variable width, some
usually 3.5–5.0 mm wide; cauline leaves 2–4, remote, c. 2–5 cm long, with lower ones often
3–4 mm wide and upper ones 2–3 mm wide, sometimes narrower (c. 1.0–1.5 mm wide). All
leaves ±acute to subobtuse; margins densely papillose-serrulate. Lower bract sometimes
remote, subherbaceous, very narrow, acute, c. 1–2 cm long, usually shorter than inflorescence;
middle bracts conspicuous, membranous, whitish straw-brown, castaneous at base,
lanceolate-aristate, usually 6–9 mm long. Inflorescence nodding (on arcuate upper part of
stem), ±dense to inconspicuously interrupted at base, spike-like, usually 1.2–2.5 × c. 1 cm, of
2–7 few-flowered clusters. Bracteoles scarious, often brownish at base, ±ciliate, ovatelanceolate,
c. 2.5 mm long. Tepals subequal, glabrous, lanceolate, acuminate, usually 2.8–3.7 mm
long, dark castaneous-brown, with narrow paler membranous margins. Stamens 6; anthers
0.9–1.2 mm long; filaments 0.3–0.4 mm long; style 0.6–0.9 mm long; stigmas c. 1.1–1.7 mm
long. Capsule ±ovoid, subacute, pale to dark castaneous-brown, ±exceeding perianth; capsule
segments 2.5–3.0 × 1.9–2.2 mm. Seeds subglobular, pale to dark brown, 1.4–1.8 mm long,
1.1–1.3 mm wide; appendage yellowish, c. 0.2 mm long, fibrillate. 2n=24, fide R.Franzén, in
A.Strid & Kit Tan, Mount. Fl. Greece 2: 742 (1991).
Southern Europe: Balkan Peninsula and S Italy. 13: ALB, BUL, GRC, ITA, YUC. Subalpine
meadows, usually on silicic rocks, usually between 1800 and 2500 m. Map 54.
13. ALBANIA: Permët, Biovizhdë, Nemerçka Mtns, A.Alston & N.Y.Sandwith 1811 (K). BULGARIA: Pirin
Mtns, Banderica, below Mt Vikhren, c. 2200 m, J.Kirschner 4465 (PRA). GREECE: Macedonia, Samarina,
Mt Smolikas, 2150 m, E.K.Balls & H.W.Gourlay 3418 (K); Thessalia, Mt Olympos, 2600–2650 m, A.Strid
1505 (C, LD); Pindhos Mtns, Trikala, Dokimi, P.E.E.Sintenis [It. Thessal.] 844 (LD, PRC); Evritania,
Karpenision, Mt Timfrestos, 2050–2100 m, L.-Å.Gustavsson 1626 (LD). ITALY: Basilicata, Mt Volturino,
1830 m, O.Gavioli s.n. (FI). CRNA GORA: Javorje Planina [Mtns], Jul 1903, J.Rohlena (PRC); Štavna, Mt.
Kom, Jul 1903, J.Rohlena (PR, LD). F.Y.R. MACEDONIA: Štrba, Šar Planina [Mtns], 2100 m, 18 Sep
1922, K.Vandas (PR).
At the sites with a common occurrence of L. pindica and L. spicata subsp. italica in the
Balkans, intermediates between the two taxa are found. They are probably of a hybrid origin.
A specimen from E Anatolia [Amasya, Mt Ak-dagh, J.F.N.Bornmüller [Pl. Exs. Anatol. Or.]
884 (WU)] has most of the attributes of typical L. pindica but the material is insufficient for
a sure identification.
29. Luzula spicata (L.) DC., in J.B.A.P. de M. de Lamarck & A.P. de Candolle, Fl.
Franç., 3rd edn, 3: 161 (1805)
Juncus spicatus L., Sp. Pl. 330 (1753); Gymnodes spicata (L.) Fourr., Ann. Soc. Linn. Lyon, sér. 2, 17: 173
(1869); Juncoides spicata (L.) Kuntze, Revis. Gen. Pl. 2: 725 (1891). T: Sweden, Lapland, C.Linnaeus 125;
lecto: Flora Lapponica herbarium, Institut de France, Paris, designated here.