Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

JUNCACEAE (Luzula)
31. Luzula caespitosa (E.Mey.) Steud., Syn. Pl. Glumac. 2: 294 (1855)
Luzula pediformis var. caespitosa [Gay ex] E.Mey., Linnaea 22: 420 (1849); Juncoides caespitosa (E.Mey.)
Kuntze, Revis. Gen. Pl. 2: 724 (1891); Luzula pediformis subsp. caespitosa (E.Mey.) Guinea, Fl. Santander
355 (1953). T: Spain, Asturias, Cangas de Tineo, Pico de Arvas, 1835, M.C.Durieu [Pl. Sel. Hispan. Lusit.]
216; lecto: K, designated here; isolecto: W.
Luzula caespitosa subsp. sanabriae P.Monts., Anales Inst. Bot. Cavanilles 21(2) [1963]: 473 (1964).
T: Spain, Zamora, Sierra Segundera, Rivadelago, Moncalvo, laguna de Lacillos, 1700–1800 m, Jun 1948,
T.M.Losa & P.Montserrat; holo: BC114621; iso: BCF4132, 46673.
Luzula caespitosa subsp. iberica P.Monts., Anales Inst. Bot. Cavanilles 21(2) [1963]: 473 (1964). T: Spain,
Burgos, Cordillera Ibérica, Laguna Negra, sobre Neila, 1800 m, P.Font Quer 339; holo: BC63341.
Map: P.Montserrat, Anales Inst. Bot. Cavanilles 21(2): map 4 (1964).
Perennials usually 20–30 cm tall, loosely caespitose, densely hairy at base; rhizome long,
±densely branched, subvertical to ascending, c. 2–3 mm in diam., often covered with greyish
brown fibrous remains of old leaf sheaths. Stem up to 1 mm in diam., upper part nodding.
Basal leaves usually ±canaliculate, usually 4–7 (–11) cm long, up to 2 mm wide, (leaves of
new tillers and intravaginal shoots much narrower, c. 0.5 mm). Cauline leaves 2–3; blade
usually 3–4 cm long, ±equalling or shorter than leaf sheaths. All leaves with ±papilloseserrulate
margins; apex acute to acuminate. Lower bract herbaceous at base, ±brownish,
1.0–2.0 (–2.5) cm long, usually exceeding inflorescence; middle bracts ovate-lanceolate,
membranous, c. 6–9 mm long, ciliate. Inflorescence ±nodding, usually 4–9 (–13)-flowered,
spike-like, usually congested, rarely with a ±remote basal cluster (or flower). Bracteoles
ovate, membranous, ciliate, c. 2.0–2.5 mm long. Tepals ±equal, entire to sparsely denticulate
at apex, usually 3.7–4.5 mm long, castaneous-brown with paler to ±membranous margins
above. Stamens 6; anthers 2.0–2.9 mm long; filaments 0.3–0.4 mm long; style 1.8–2.5 mm
long; stigmas 2.0–2.5 mm long. Capsule straw-brown to pale castaneous-brown, subglobose,
subacute, ±mucronate; capsule segments 3.0–3.5 × 2.6–3.0 mm long. Seeds pale brown,
broadly ovoid, usually 1.6–1.9 mm long (including a small paler apex c. 1.5 mm wide); basal
appendage yellowish, c. 0.2 mm long, fibrillate. 2n=12 [12AL], fide C.Garcia-Herran, in litt.
Mountains of NW and C Spain and EC Portugal. 12: POR, SPA. Rocky slopes and dry
mountain grasslands, usually above 1600 m. Map 61.
12. PORTUGAL: Beira Baixa, Serra de Estrela, Rua dos Mercadores, A.Fernandes, F.Sousa & J.Matos 4483
(UPS); Serra de Estrela, Torre, Manteigas, 13–25 Jun 1987, A.Polatschek (W). SPAIN: Asturias,
Leitariegos, Cueto de Arvas, P.Font Quer & W.Rothmaler [Fl. Iber. Sel.] 205 (BCF, S, UPS); Asturias, Pico
de Arvas, Jul 1898, M.Gandoger (W); Palencia, Pico de Curavacas, 16 Jul 1990, C.Garcia-Herran (NEU90-
1038; 2n=12); León, Picos del Mampodre, Cervunal de Valverde, 1800–1900 m, 5 Aug 1952, T.M.Losa &
P.Montserrat (BCF4129); León, Hayedo del Gildar, 1900 m, Garcia-Gonzales (JACA109685);
Madrid/Segovia, Sierra de Guadarrama, Siete Picos, Cercedilla – La Granja, 2080–2120 m, V.J.Aran &
M.J.Toha (JACA172898, MA); Soria, Laguna Negra, 21 Jul 1991, C.Garcia-Herran (NEU91-1089; 2n=12);
Soria, Coraleda, Pico de Urbión, 2000–2200 m, J.L.Benito (JACA19895).
A species variable in the size of its vegetative parts and in the shape and size of its
inflorescence. The variation shows considerable overlaps between local populations, and
extremes are scattered throughout the species’ range. Thus, the aberrant plants described by
P.Montserrat as subspecies are here subsumed within the species.
32. Luzula ecuadoriensis Balslev, Fl. Ecuador 11: 11 (1979)
T: Ecuador, Cotopaxi, 3550 m, 1939, E.Asplund 6494; holo: S; iso: S.
Illustration: H.Balslev, Fl. Neotrop. Monogr. 68: 62, fig. 31 (1996).
Map: H.Balslev, Fl. Neotrop. Monogr. 68: 63, fig. 32 (1996).
Perennials 50–75 cm tall, sparsely hairy to subglabrous; rhizome ascending. Basal leaves
(5–) 10–20 (–25) cm long, usually 6–10 mm wide; cauline leaves usually 3–5, with sheath to
7 cm long, usually 9–15 cm long, up to 10 mm wide; all leaves acute with densely papilloseserrulate
margins. Lower bracts similar to leaves, up to 17 cm long, distally gradually shorter
(lower inflorescence branches often in axils of upper or middle cauline leaves). Inflorescence
an interrupted ±drooping panicle in upper 1/3 (– 1/2) of plant height, composed of dense
oblong-ovoid partial inflorescences on long primary peduncles; further composed of
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