34 SPECIES PLANTARUM — FLORA OF THE WORLD (2002) Figure 8. <strong>Luzula</strong> sylvatica subsp. sylvatica. A, habit; B, flower with unripe capsule; C, capsule; D, seed; E, tepal; F, stem T.S.; G, leaf T.S. (A–G, B.Błocki, Jul 1882, KRA). Scale bars: A = 5 cm; B–F = 2 mm; G = 1 mm. Drawn by T.Szynal. Reproduced with permission, from T.Szynal & J.Mądalski, Atlas Fl. Pol. 2/2: 153 (1931).
<strong>JUNCACEAE</strong> (<strong>Luzula</strong>) <strong>Luzula</strong> sylvatica var. paularensis Pau, Bol. Soc. Aragonesa Ci. Nat. 1916: 158 (1916). T: Spain, Madrid, Sierra de Guadarrama, El Paular, C.Vicioso; syn: MA 19634; 25 Aug 1912; C.Vicioso & Beltrán; syn: MA 19641. <strong>Luzula</strong> sylvatica subsp. cantabrica P. Monts., Anales Inst. Bot. Cavanilles 21(2) [1963]: 441 (1964). T: Spain, Puer<strong>to</strong> Palombera, Reinosa, 1350 m, 26 Jun 1926, P.Font Quer; holo: BC. Plants usually (20–) 60–80 cm tall. Basal leaves (5–) 8–15 (–19) mm wide, gradually narrowed in<strong>to</strong> long acute apex; margins ciliate. Inflorescence very lax; primary branches often elongated; secondary anthelas usually ±contracted, with 30–45 flowers; terminal clusters (1–) 2–3-flowered. Outer tepals usually (1.7–) 2.3–2.7 (–3.3) mm long. Anthers (0.7–) 0.9–1.8 (–2.1) mm long; filaments usually 0.3–0.6 mm long; style 1.9–2.2 mm long; stigmas 2.2–2.6 mm long. Capsule usually ±longer than perianth, ±ovoid, usually 1.6–2.6 mm long, mucro (0.2–) 0.4–0.7 mm long. Seeds usually 1.1–1.6 mm long, c. 0.6–0.7 mm wide; appendage c. 0.1–0.2 mm long. 2n=12, fide C.Garcia-Herran (see list of specimens); H.Nordenskiöld, Hereditas 37: 328–329 (1951). Confined <strong>to</strong> the northern part of the Iberian Peninsula. 12: POR, SPA. Found in woodlands, particularly along rivers, on damp soils. Map 28. 12. PORTUGAL: Serra do Gerês, Albergaria, 700 m, C.Garcia-Herran 91-1134 (NEU; 2n=12); Serra da Estrela, Poço do Inferno, J.Ma<strong>to</strong>s, Cardoso & A.Marques 7905 (PR); Serra da Estrela: Lagoa do Covao do Curral, A.Fernandes, R.Fernandes & F.Sousa 4521 (LD); Al<strong>to</strong> Douro, Bragança, Pena de Águia, 750 m, P.Silva, B.Rainha & J.Martins 7788 (LD); Cinfaes, Fonte Nova, J.Paiva, J.Ma<strong>to</strong>s & A.Marques 8481 (PR). SPAIN: Asturias, Grado, San Pelayo, 2 May 1978, J.J.Lastra (FCO 24033); Asturias Luarca, along Esva R., near Pontigón, 24 Mar 1974, T.E.Díaz (FCO 23640); Salamanca, Miranda del Castañar, 690 m, 4 Apr 1985, D.Sánchez-Mata & V. de la Fuente (FCO 17362). A variable subspecies with several local variants. The eastern populations approach the type subspecies. 10c. <strong>Luzula</strong> sylvatica subsp. sicula (Parl.) K.Richt., Pl. Eur. 1: 183 (1890) <strong>Luzula</strong> sicula Parl., Nuov. Gen. Sp. Monocot. 59 (1854); <strong>Luzula</strong> maxima subsp. sicula (Parl.) Nym., Consp. Fl. Eur. 751 (1882); <strong>Luzula</strong> sieberi subsp. sicula (Parl.) Pignatti, Fl. Ital. 3: 444 (1982), nom. inval. T: Sicily: ‘monti di Sicilia nelle Madonie, in Montesoro, nei boschi di Caronia, di Mistretta, di Cannata, di Francavilla’; syn: n.v. Plants usually 30–45 (–65) cm tall. Basal leaves usually 3–4 (–6) mm wide, with elongated acuminate tip; margins and base very densely ciliate. Inflorescence lax, few-flowered; secondary anthela usually with (5–) 8–13 (–16) flowers; terminal clusters (1–) 2–3 (–4)flowered. Outer tepals usually 4.3–4.8 (–5.2) mm long. Anthers (1.7–) 2.2–2.5 (–3.0) mm long; filaments 0.2–0.6 mm long; style 1.7–1.9 mm long; stigmas on average 2.4–2.6 mm long. Capsule usually ±equalling perianth or shorter, ±ovoid, 3.5–4.1 (–4.4) mm long; mucro (0.2–) 0.4–0.8 mm long. Seeds (1.5–) 1.7–2.0 (–2.2) mm long, 1.1–1.3 mm wide; appendage c. 0.2 mm long. 2n=12, fide C.Garcia-Herran (see list of specimens). Southernmost Italy and Sicily. 13: ITA, SIC. In mountains, on volcanic slopes with sparse vegetation or broad-leaved (mostly Fagus sylvatica) woodlands. Map 29. 13. ITALY: Calabria, E ridge of Montenero, 1400–1600 m, S.Snogerup & B.Snogerup 2081 (LD). SICILY: Messina, Floresta, 1350 m, Kraehembuhl 91-684 (NEU; 2n=12); Madonie, 600–1000 m, 18 Jul 1874, G.Strobl (K); Messina, Floresta, Mt Pojummoru, 1300 m, R.K.Brummitt, Hunt & Leistner 5104 (K); Valdemone, A.Todaro [Fl. Sic. Exs.] 649 (K, W); E and NE slopes of Mt Etna, 2250 m, 19–20 Aug 1873, G.Strobl (WU). 10d. <strong>Luzula</strong> sylvatica subsp. sieberi (Tausch) K.Richt., Pl. Eur. 1: 183 (1890) <strong>Luzula</strong> sieberi Tausch, Flora 19: 423 (1836); <strong>Luzula</strong> maxima β [unranked] angustifolia Čelak., Prodr. Fl. Böhmen 1: 84 (1867), nom. illeg., non Roth (1788); <strong>Luzula</strong> sylvatica var. sieberi (Tausch) Buchenau, Bot. Jahrb. Syst. 12: 92 (1890); <strong>Luzula</strong> maxima subsp. sieberii (Tausch) P.Fourn., Fl. Compl. Plaine Franç. 527 (1928). T: [Austrian Alps] F.W.Sieber [Herb. Fl. Austr.] 104, ut Juncus maximus β angustifolius; lec<strong>to</strong>: PRC, designated here; isolec<strong>to</strong>: BRNM. Illustration: S.Pignatti, Fl. Ital. 3: 444 (1982). 35
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