Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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40 SPECIES PLANTARUM — FLORA OF THE WORLD (2002) 4 Tepals stramineous to pale brownish, with apex often ±reflexed; inflorescence very lax, with rigidly divaricate branches 17. L. divaricata 4: Tepals blackish brown to dark castaneous-brown, with apex not reflexed; inflorescence lax to subdense, ±nodding to suberect, with branches not rigidly divaricate 5 Basal leaves densely to sparsely ciliate. [Mexico to Panama] 14. L. denticulata 5: Basal leaves subglabrous. [Outside C America] 6 Capsule segments usually 3.0–3.5 × 1.8–2.0 mm 19. L. hitchcockii 6: Capsule segments to 2.9 mm long and to 1.6 mm wide 7 Anthers ±equalling filaments or shorter; style up to 0.5 mm long 8 Plants usually more than 30 cm tall; cauline leaves (3–) 4–5, usually 5–10 mm wide 22. L. parviflora 8: Plants usually up to 30 cm tall; cauline leaves 1–3, usually less than 5 mm wide 9 Cauline leaves bluish green to greyish green, (2–) 3; lower bract 1.5–2.5 cm long 18. L. piperi 9: Cauline leaves mid-green, 1–2; lower bract to 1 cm long 15. L. wahlenbergii 7: Anthers (2–) 3–5 (–9) times longer than filaments; style usually longer than 0.5 mm 10 Capsule segments to 2.4 mm long; seeds stramineous to pale brown 10: Capsule segments 2.4–3.0 mm long; seeds brown to dark brown 11 Cauline leaves 10–20 cm long, usually 3–6 mm wide; anthers 24. L. alpinopilosa usually 3–4 times longer than filaments 11: Cauline leaves up to 6 cm long, usually 4.5–8.0 mm wide; 21. L. desvauxii anthers usually 5–7 times longer than filaments 20. L. glabrata 13. Luzula gigantea Desv., J. Bot. (Desvaux) 1: 145 (1808) Juncoides spadicea Kuntze var. gigantea (Desv.) Kuntze, Revis. Gen. Pl. 2: 724 (1891); Juncoides gigantea (Desv.) E.Sheld., Minnesota Bot. Stud. 1: 62 (1894). T: Colombia, in montibus humidis Quindio in regno Santa-fe de Bogota, A.Humboldt & A.J.Bonpland; holo: P. Luzula paniculata Desv., J. Bot. (Desvaux) 1: 147 (1808). T: ‘In America calidiore’; holo: P. Illustrations: M.Barros, Darwiniana 10: 318, fig. 10 (1953); H.Balslev, Fl. Neotrop. Monogr. 68: 53, fig. 26 (1996). Map: H.Balslev, Fl. Neotrop. Monogr. 68: 54, fig. 27 (1996). Perennials 20–90 cm tall; rhizome ascending, 1.5–3.0 mm in diam.; stolons long, ascending. Basal leaves 10–30 cm long, (5–) 10–20 mm wide, flat; margin densely papillose-serrulate, ciliate in young leaves; cauline leaves usually 2–4, with upper ones usually up to 10–13 mm wide; sheaths to 6 cm long; leaf tip acute. Inflorescence a rich loose compound panicle constituting more than one third of plant height, storied and subdivided by 5–15 cm long rhachis segments into 4–5 subequal nodding anthelate parts; flowers borne singly (rarely in clusters of 2–3). Bracts of primary inflorescence branches resembling cauline leaves, with proximal ones to 20 cm long and distal ones gradually shorter; bracts on distal branches brownish-membranous, linear-lanceolate, c. 1.0–1.5 cm long. Bracteoles ovate to ovatelanceolate, brownish-membranous, 1.6–2.0 mm long, usually entire, glabrous or sparsely ciliate, inserted 0.2–0.5 mm below flowers. Tepals ±unequal, glabrous, acuminate, usually dark castaneous brown, margins paler; outer tepals usually 2.2–2.7 mm long; inner tepals usually 2.0–2.5 mm long. Stamens 6, 1.0–1.5 mm long; anthers 0.3–0.6 mm long; filaments 0.7–0.9 mm long; style 0.2–0.3 mm long; stigmas ±patent, 0.6–0.8 mm long. Capsule broadly ellipsoidal, apiculate, shorter than perianth, castaneous, shining; capsule segments often slightly
JUNCACEAE (Luzula) mucronate, 1.6–1.9 × c. 1.1 mm long. Seeds ±ellipsoidal, (0.9–) 1.0–1.1 × 0.5–0.6 mm; outer seed-coat white to light brown; appendage almost not visible, c. 0.1 mm long; base fibrillate. Fig. 9. South America, in the Andes from Venezuela and Colombia to Bolivia and N Argentina. 82: VEN. 83: BOL, CLM, ECU, PER. 85: AGW. Open mountain slopes, páramo, along streams, usually between 3000 and 4000 m. Map 33. 82. VENEZUELA: Mérida, Trujillo, J.Linden 412 (BM, BR, G, P, US). 83. BOLIVIA: Cochabamba, Chapare road at km 110, 3000 m, R.F.Steinbach 611 (LPB, NY, S, US, WIS); La Paz, Larecaja, near Sorata and Mt Sanquiarie, 3200 m, G.Mandon 1423 (BM, K, NY, P, S, US). COLOMBIA: Cauca, Cordillera Central, Puracé, 3450 m, K.Sneidern 1740 (S); Boyaca, Sierra Nevada de Cocuy, Quebrada de las Playas, 4100 m, P.J.Grubb et al. 321 (K, US). ECUADOR: Imbabura, Lago San Marcos Cayambe, Wash et al. 5460 (K); Imbabura, Mojanda, 10 km SSW Otavalo, 2900–3150 m, B.Sparre 13476 (S); Pichincha, Mt Corazón, E.Asplund 9684 (S). PERU: Huacás, Santiago de Chuco, 2800 m, A.Sagástegui, J.Mostacero & Diestra 11913 (AAU, MO, K, NY, F); Mito, c. 3000 m, J.F.Macbride 3367 (S). At first sight very similar to SE Asian Luzula effusa (the only other taxon having storied inflorescence) but differs from it in having smooth capsules, longer outer tepals, anthers substantially shorter than filaments, narrower seeds and longer bracteoles. 14. Luzula denticulata Liebm., Vidensk. Meddel. Dansk Naturhist. Foren. Kjøbenhavn 2: 46 (1850) Luzula parviflora var. denticulata (Liebm.) Buchenau, Bot. Jahrb. Syst. 7: 171 (1885). T: Mexico, Orizaba, Vaqueria del Jacal, 3050 m, F.Liebmann; lecto: C, fide L.Hämet-Ahti, Ann. Bot. Fenn. 8: 370 (1971); isolecto: C, G, HAL, K, P, S, UPS. Luzula laetevirens Liebm., Vidensk. Meddel. Dansk Naturhist. Foren. Kjøbenhavn 2: 46 (1850). T: Mexico, Orizaba, 2750–3050 m, F.Liebmann; lecto: C, fide L.Hämet-Ahti, Ann. Bot. Fenn. 8: 370 (1971); isolecto: C, P. Luzula latifolia Liebm., Vidensk. Meddel. Dansk Naturhist. Foren. Kjøbenhavn 2: 47 (1850). T: Mexico, Oaxaca, Cumbre de Ixtepec, 3050 m, F.Liebmann; lecto: C, fide L.Hämet-Ahti, Ann. Bot. Fenn. 8: 370 (1971); isolecto: C. Luzula gigantea var. vulcanica Woodson, in R.E.Woodson & R.J.Seibert, Ann. Missouri Bot. Gard. 26: 278 (1939). T: Panama, Chiriquí, Loma Larga, Volcán Chiriquí, 2500–3380 m, R.E.Woodson jr., P.H.Allen & R.J.Seibert 1094; holo: MO; iso: GH, NY, US, photo AAU. Illustration: H.Balslev, Fl. Neotrop. Monogr. 68: 56, fig. 28 (1996). Map: H.Balslev, Fl. Neotrop. Monogr. 68: 54, fig. 27 (1996). Perennials 15–80 cm tall; rhizome ascending, long, often with long ascending stolons. Basal leaves 5–25 cm long, 8–12 mm wide; cauline leaves usually 3–5 (–8), usually 6–8 cm long, 4–5 mm wide. All leaves usually densely ciliate, less often subglabrous, acute, with ±papilloseserrulate margins. Lower bract herbaceous, up to 6 cm long; next upper bract linear, brownish membranous, ciliate, c. 1 cm long. Inflorescence terminal, a loose nodding anthelately arranged panicle 5–20 × 3–10 cm; flowers borne singly. Bracteoles brownish-membranous, ±ovate, acute, 1.2–1.6 mm long, ciliate, inserted c. 0.1–0.2 mm below flower. Tepals subequal, glabrous, narrowly-lanceolate, acuminate, pale to dark brown, 1.7–2.4 mm long. Stamens 6; anthers 0.4–0.7 mm long, equalling or longer than filaments; filaments 0.3–0.5 mm long; style 0.1–0.2 mm long; stigmas 0.8–1.0 mm long. Capsule dark brown, shining, ovoid, acute, often equalling or exceeding perianth; capsule segments 1.5–1.7 × 1.0–1.3 mm. Seeds oblong-ellipsoidal, brown, 1.0–1.1 mm long, 0.6–0.7 mm wide; appendage indistinct, to c. 0.1 mm, fibrillate. Fig. 10. Confined to Central America, from C Mexico to Panama. 79: MXC, MXE, MXG, MXS, MXT. 80: COS, GUA, PAN. Grasslands, pastures and open woodlands, between 200–3900 m. Map 34. 79. MEXICO CENTRAL: Distrito Federal, Contreras, Los Dinamos, 2800 m, A.Ventura 3580 (AAU); Distrito Federal, Mt Ajusco, 3840 m, J.H.Beaman 2806 (DS, MICH, US); México State, La Ciénega, Iztaccíhuatl, R.Galván 735 (AAU); Puebla, Sierra Negra, Mt Orizaba, 3200 m, E.K.Balls et al. 4475 (K, UC, US). MEXICO NORTHEAST: Hidalgo, Epazoyucan, Nopalillo, 2850 m, R.Galván 580 (AAU); Hidalgo, El Chico, E.Lyonnet 265 (CAS, GH, MEXU, MO, NY, K, US). 80. COSTA RICA: Cartago, Volcán Irazú, 3000 m, K.Barringer, Christenson & L.D.Gomez 3273 (AAU); Cartago, Cerro de la Muerte, 3000–3500 m, 41
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Page 3 and 4: CONTENTS Contributors iv Species Pl
Page 5 and 6: Species Plantarum Steering Committe
Page 7: PREFACE This volume is the first of
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Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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