Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

JUNCACEAE (Luzula)
Luzula campestris A.II.a.2. [unranked] luteola Asch. & Graebn., Syn. Mitteleur. Fl. 2: 522 (1904); Luzula
campestris var. luteola (Asch. & Graebner) V.I.Krecz., Žurn. Russk. Bot. Obshch. Akad. Nauk S.S.S.R. 12: 490
(1928). T: not designated.
Luzula campestris var. insularis Briq., Prodr. Fl. Corse 1: 246 (1910); Luzula campestris proles insularis
(Briq.) Rouy, Fl. France 13: 265 (1912); Luzula insularis (Briq.) Prain, Ind. Kew., suppl. 5: 157 (1921).
T: Corse, entre Ste. Lucie et La Trinité, 50 m, 7 May 1907, J.I.Briquet; lecto: G, designated here.
Luzula chabertii Rouy, Fl. France 13: 266 (1912); Luzula campestroides H.Lév., Bull. Acad. Int. Géogr. Bot.,
ser. 4, 25: 48 (1917), nom. illeg. T: [France] Forêt d’Fontainebleau dans les Ventes Bourbon, 16 May 1871,
A.C.Chabert; lecto: FI, fide J.Kirschner, Taxon 39: 112 (1990).
Luzula campestris var. rectirama Murr, Neue Übers. Bl-Pfl. Vorarlberg 46 (1923). T: [Austria] Nofels,
Gisingen und Letzebühel b. Feldkirch, J.Murr; syn: n.v.
Luzula campestris var. terana P.Monts., Anales Inst. Bot. Cavanilles 21(2): 492 (1964). T: Zamora,
Sanabria, Tera R., Furnio de Lacillos, 1600 m, Jun 1945, T.M.Losa; holo: BCF.
Luzula campestris subsp. iberica P.Monts., Anales Inst. Bot. Cavanilles 21(2): 492 (1964). T: Madrid,
Sierra de Guadarrama, Peñalara, 25 May 1912, C.Vicioso; holo: BC63308.
Illustrations: H.E.Hess, E.Landolt & R.Hirzel, Fl. Schweiz 1: 521 (1967); J.Kirschner, Folia Geobot. Phytotax.
28: 153, fig. 3 (1993).
Map: H.Meusel, E.Jäger & E.Weinert, Vergl. Chorol. Zentraleur. Fl. 1: 88 (1965).
Perennial, 5–25 cm tall, ±densely ciliate, with creeping rhizome and short stolons. Stem with
numerous basal and 2–3 cauline ±flat leaves; upper leaves 2–4 cm × 1.7–3.3 mm, with
sparsely papillose-serrulate margins. Clusters 3–9-flowered; peduncles erect, often ±flexuose,
usually at least one nodding. Lower bract 1–3 cm long, shorter than inflorescence. Bracteoles
2.0–2.5 mm long, whitish-membranous, ciliate-fimbriate. Tepals 2.8–4.1 mm long,
castaneous brown to dark brown. Anthers 1.3–1.8 mm long, 3.0–4.5 times longer than the
0.3–0.6 mm long filaments; style 1.0–1.5 mm long; stigma 1.8–3.0 mm long, ±persistent.
Capsule ±equalling tepals; capsule segments 2.1–3.0 mm long, dark brown. Seeds
subglobular, 1.0–1.1 × 0.8–1.0 mm; basal appendage 0.4–0.7 mm long. Mean length of
middle stomata in upper cauline leaves 30–35 µm. 2n=12 [12AL], fide H.Nordenskiöld,
Hereditas 37: 336–337 (1951); J.Kirschner, Pl. Syst. Evol. 200: 5 (1996). Fig. 22.
Europe, except for the eastern part, and North Africa. Introduced to North America, New
Zealand, Australia, Macaronesia, Falkland Is. 10: DEN, FIN, FOR, GRB, IRE, NOR, SWE.
11: All. 12: BAL, COR, FRA, POR, SAR, SPA. 13: ALB, BUL, GRC, ITA, ROM, TUE,
YUC. 14: BLR, BLT, RUC, RUN, RUW, UKR. 20: ALG, MOR. 21: azo, mdr. 50: nsw, tas,
vic. 51: nzn, nzs. 72: nfl. 75: mas. 90: fal. Map. 87 [secondary occurrence not shown]
11. GERMANY: Berlin, Jungfernhaide, 120 m, Lackowitz [A.Kneucker, Cyp. Junc. Exs.] 60 (GZU, PR).
POLAND: Chrzanów, Płaza, A.Jasiewicz & Pałkowa [Fl. Polon. Exs.] 390 (WRSL). SWITZERLAND:
Chur, Campodels, C.Coaz [J.Braun-Blanquet, Fl. Raet. Exs.] 611 (BC, G, GZU). 12. FRANCE: Seine-et-
Oise, Forêt de Sénart, De Baire [Duffour, Soc. Franc.] 2242 (BC, P). PORTUGAL: Algarve, Sierra
Monchique, 600 m, P.Montserrat et al. (JACA590668). SPAIN: Madrid, El Escorial de Guadarrama,
15 May 1966, M.Mayor (MAF87300); Burgos, Lunada, Portillo de Lunada, 1500–1600 m, P.Montserrat
(JACA173783). 14. BELORUSSIYA: Minsk, 12 May 1986, Tzvelev (LE). RUSSIA NORTHWEST:
Pskov, V.Andreev 1683 (LE). 90. FALKLAND IS.: East Falklands, Port Stanley, Beatrice Cove, 10 Nov
1945, Hamilton 12 (BM).
Numerous literature records under the name L. campestris from outside Europe mostly refer
to other species in the sect. Luzula. In Spain and Portugal, subsp. campestris is rather
variable, particularly in the flower size, inflorescence appearance and length of stolons; in
other regions, the variation is very limited.
56b. Luzula campestris subsp. nevadensis P.Monts., Anales Inst. Bot. Cavanilles
21(2): 492 (1964)
T: Spain, Sierra Nevada, Puerto Lobo, 2100 m, 14 Jul 1923, P.Font Quer; holo: BC150865.
Map: P.Montserrat, Anales Inst. Bot. Cavanilles 21(2): map 8 (1964).
Perennial, usually 20–32 cm tall; rhizome long, creeping; stolons present. Basal leaves
sparsely ciliate, c. 3.0–4.5 mm wide; cauline leaves 2–3, with upper ones 3.5–5.0 cm long,
2.0–2.8 mm wide. Lower bract usually 1.2–2.0 cm long, shorter than inflorescence. Clusters
105