Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

JUNCACEAE (Luzula)
with sparse fibrous cover. Basal leaves usually (3–) 4–7 (–9) cm long, (3–) 4–5 (–6) mm
wide; cauline leaves usually 3, usually 2–4 (–6) cm long. All leaves subacute to subobtuse;
margins distinctly papillose-serrulate. Lower bract ±brownish-membranous, 1.0–1.5 (–2.0) cm
long, shorter than inflorescence. Inflorescence decompound, corymbiform; main branches
thin, straight to arcuate; lower branches often patent, sometimes erect, each with dense
glomeruliform (5) 10–15 (25)–flowered cluster, with one cluster often larger, subsessile.
Bracteoles ovate, c. 1.5 mm long, transparent-scarious, ±ciliate. Tepals unequal, ±oblonglanceolate,
glabrous, straw-yellow, becoming ±dirty yellow, with broad membranous
margins; inner tepals 2.8–3.2 mm long; outer tepals shorter, c. 2.5–2.7 mm long. Stamens 6;
anthers 1.1–1.8 mm long; filaments 0.3–1.1 mm long, distinctly elongating during flowering
(anther/filament ratio thus changing); style (0.9–) 1.0–1.3 mm long; stigmas 0.5–0.8 mm
long. Capsule oblong-ovoid, acuminate, mucronate, deep castaneous-brown, shining; capsule
segments 2.4–2.8 × 1.4–1.5 mm long including a mucro 0.2–0.3 mm long. Seeds oblongovoid,
paler brown, 1.0–1.3 mm long including a paler apex c. 0.2 mm long and 0.7–0.8 mm
wide; appendage c. 0.1 mm, fibrillate. 2n=12, fide H.Nordenskiöld, Hereditas 37: 328–329
(1951).
Southwestern part of the Alps, E Pyrenees and N Italian mountains. 11: AUT, SWI. 12: FRA,
SPA. 13: ITA. Growing in alpine grasslands and screes, usually on siliceous substrates, from
c. 1700 m. Map 24.
11. AUSTRIA: Nordtirol, Gurgler Tal, 1900–2200 m, 13 Sep 1989, A.Polatschek 17612 (W); Oetztal,
Klebealpe, 2000 m, 11 Jun 1937, Fischer (K). SWITZERLAND: Valais, Saas–Fée, 2100 m, L.Marret [Exs.
Fl. Valais] 296 (S); Valais, Alpe de la Barma, 2200 m, F.O.Wolf [A.Kneucker, Cyper. Junc. Exs.] 21 (PR);
Graubünden, Ober Engadin, Bernina Pass, 2330 m, O.Hedberg 4022 (UPS). 12. FRANCE: Haute-Savoie,
Massif du Mont Buet, Val de Bérano, 2000 m, B. de Retz 40427 (W); Hautes-Alpes, Goudran, Montgenèvre,
2000–2200 m, C.Magnier [Fl. Select. Exs.] 1823 (LD). ANDORRA: Tristaina Lakes, 2800 m, Lewis 485 (K).
SPAIN: Setcasas, Mt Morens, E.Bourgeau [Pyren. Esp.] 277 (K). 13. ITALY: Levico, Pancrotto, Jul 1926,
Meebold (K).
A species with limited variation. The westernmost populations were recognized as subsp.
pyrenaica on the basis of the inflorescence habit, broader leaves, longer tepals and smaller
seeds. However, despite the fact that eastern populations show a lower frequency of broadleaved
individuals, the other characters do not exhibit any geographical tendency, and the
extremes are often found within populations throughout the geographical range of the species.
9. Luzula luzuloides (Lam.) Dandy & E.Willm., J. Bot. 76: 352 (1938)
Juncus luzuloides Lam., Encycl. 3: 272 (1789). T: France: ‘Lorraine Allemande, près des rives de la Sarre’,
de l’Aumont; lecto: P-LAM, fide S.Snogerup, in P.H.Davis, Fl. Turkey 9: 28 (1986).
Luzula albida var. macrantha Beck, Fl. Nieder-Österreich 1: 159 (1890); Luzula nemorosa A.I. [unranked]
macrantha (Beck) Asch. & Graebn., Syn. Mitteleur. Fl. 2(2): 502 (1904); Luzula albida subvar. macrantha
(Beck) Rouy, Fl. France 13: 261 (1912). T: Austria, Mt Schneeberg, G.Beck; holo: n.v.
Luzula albida var. parviflora Döll, Fl. Baden 1: 323 (1857); Luzula nemorosa var. parviflora (Döll) Asch. &
Graebn., Syn. Mitteleur. Fl. 2(2): 504 (1904); Luzula luzuloides var. parviflora (Döll) I.Grint., Fl. Rep. Soc.
Roman. 11: 591 (1966). T: [Germany] Baden, Untergrombach, Döll; syn: n.v.
Luzula nemorosa B.II. [unranked] hylogiton Asch. & Graebn., Syn. Mitteleur. Fl. 2(2): 504 (1904).
T: Germany, Harz; n.v.
Maps: H.Meusel, E.Jäger & E.Weinert, Vergl. Chorol. Zentraleur. Fl. 1: 87 (1965); E.Hultén & M.Fries, Atlas
N. Eur. Vasc. Pl. 1: map 165 (1986).
Perennials usually 35–75 cm tall, loosely caespitose to caespitose, densely ciliate; rhizome
horizontal to ascending, branched; stolons usually present, ascending. Basal leaves
numerous, usually 12–18 (–30) cm long, width variable but ranging from (3–) 4–5 (–6) mm;
cauline leaves usually 4–5, to c. 25 cm long, with the upper ones often overtopping
inflorescence. All leaves with acute tip; margins papillose-serrulate. Lower bracts herbaceous,
of variable length, usually overtopping inflorescence, less often shorter. Inflorescence rich,
decompound, ±anthelate; main branches usually numerous, with secondary branches each
bearing ±dense or lax anthela of (2–) 3–6 (–8)-flowered clusters. Bracteoles ±ovate,
subglabrous to ciliate, usually 1.0–1.5 mm long. Tepals glabrous, subequal with inner longer
by c. 0.5 mm, ±lanceolate, (2.0–) 2.3–3.8 (–4.2) mm long. Stamens 6; anthers 1.0–1.8 mm
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