Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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52 SPECIES PLANTARUM — FLORA OF THE WORLD (2002) Perennials (20–) 40–80 (–100) cm tall, loosely caespitose, subglabrous; rhizome creeping or ascending, branched; stolons usually present, short or to c. 10 cm long. Leaves with densely papillose-serrulate margins; apex acute; basal leaves few; cauline leaves 3–6 (upper 2 or 3 represent leaf-like bracts). Inflorescence usually lax, usually constituting 1/3– 1/2 of plant height, decompound, storied; partial inflorescences subtended by leaf-like bracts; branches diverging at wide angles; flowers borne singly, rarely in 2 (–3)-flowered clusters. Tepals equal, glabrous, ±lanceolate, usually 1.8–2.8 mm long, pale reddish brown to dark castaneous-brown. Stamens 6; anthers 0.5–1.1 mm long; filaments 0.6–1.0 mm long; style usually 0.4–0.5 mm long; stigmas 1.0–1.5 mm long. Capsule obovoid to ±ellipsoid, subacute to acute, with mucro 0.2–0.4 mm long, distinctly papillose above when ripe, equalling or ±longer than perianth; capsule segments c. 2.1–2.3 × 1.1–1.4 mm. Seeds deep brown, 1.2–1.4 mm long, (0.6–) 0.7–0.8 mm wide; apex usually distinct; basal appendage indistinct, to 0.1 mm, fibrillate. A widely distributed variable complex of populations extending from E Nepal to SW China, Taiwan and S Malesia. The variation is centred in Sichuan and adjacent territories where two varieties linked by occasional intermediates are recognized. The whole group is in need of revision. Most common habitats include humid mountain forests, wet scrub, shady ravines and cliff-faces. Lower cauline leaves usually 4–11 mm wide; tepals ±brown to deep (castaneous-)brown; stem usually less than 1.5–2.0 mm in diam. 23a. var. effusa Lower cauline leaves usually 10–21 mm wide; tepals paler reddish brown to brown; stem usually 3–6 mm in diam. 23b. var. chinensis 23a. Luzula effusa Buchenau var. effusa Illustrations: Muh-Tsuen Kao & C.E.Devol, in Hui-Lin Li, Fl. Taiwan 5: 155, plate 1304 (1978); H.J.Noltie, Fl. Bhutan 3(1): 263, fig. 25 (1994); K.F.Wu, Fl. Reipubl. Popularis Sin. 13(3): 240, plate 58 (1997). Plants usually to (50–) 80 cm tall, often slender; stolons short to long. Stem usually 1–2 mm diam. Cauline leaves usually 4–11 mm wide. Tepals ±brown to deep (castaneous-)brown, usually 1.8–2.8 mm long. Anthers 0.5–1.1 mm long; filaments usually 0.6–0.8 (–1.0) mm long. The East Himalaya from E Nepal to Manipur, N Burma, SW and C China, Taiwan, Luzon, Borneo, Sulawesi and New Guinea. 36: CHC, CHN, CHT. 38: TAI. 40: EHM, NEP. 41: BMA. 42: BOR, NWG, PHI, SUL. Humid mountain woodlands, shrubby slopes and subalpine ravines. Map 45. 36. TIBET: Kongbo, Doshong La, 4000 m, F.Ludlow, G.Sherriff & G.Taylor 4676a (BM, UPS). CHINA SOUTH-CENTRAL: Sichuan, Mt Omei [Emei], Chiu-lo Tung, 2000 m, F.T.Wang 23312 (S). CHINA NORTH-CENTRAL: Gansu ['Kansu orient.'], 4 Sep 1890, Potanin (K, LE). 38. TAIWAN: Taichung, W slope of Mt Loyehwei-shan, Shimizu & Chuang 20183 (S). 40. NEPAL: Tamur Valley, Topke Gola, c. 3530 m, J.D.A.Stainton 976 (UPS). BHUTAN: Pele-La, Gould 345 (K). 42. BORNEO: Sabah, Kinabalu, Masilau R., M.S.Clemens 51371 (K). NEW GUINEA: Papua, Goilala, Mt Albert Edward, 3600 m, J.R.Croft et al. 61394 (BM, K). PHILIPPINES: Luzon, Mt Pulog, E.D.Merrill 6490 (K). SULAWESI: Enrekang, Rante Mario, 3300 m, P.J.Eyma 879 (K). A very variable taxon. Rarely, very young plants have the inflorescence only indistinctly storied. Plants from Taiwan and Malaya have tepals usually 2.5–2.8 mm long, longer than capsules. The Malayan plants have more numerous basal leaves and, on average, longer stolons. Intermediates between the two varieties are not rare. 23b. Luzula effusa var. chinensis (N.E.Br.) K.F.Wu, J. E. China Norm. Univ., Nat. Sci. Ed., 1992(3): 92 (1992) Luzula chinensis N.E.Br., J. Linn. Soc., Bot. 36: 161 (1903). T: China, [Sichuan] Szechuan, A.Henry 8829; syn: K; S (fragment); China, Szechuan, near Tibetan frontier, Tachienlu [Kangding], A.Pratt 360; syn: BM, K. Plants usually 50–100 cm tall, robust; stolons short. Stem usually 2–6 mm in diam. Cauline leaves usually to 20 cm long and 10–21 mm wide. Tepals usually 1.9–2.3 mm long, paler reddish brown to brown. Anthers 0.5–0.6 mm long; filaments 0.6–0.8 mm long.
JUNCACEAE (Luzula) Most frequently found in Sichuan and Yunnan, less common in Tibet, Hubei, Guizhou and N Burma. 36: CHC, CHT. 41: BMA. Mesic to humid woodlands. Map 46. 36. CHINA SOUTH-CENTRAL: Hubei [originally Hupeh, without detailed locality], A.Henry 6809 (K); Sichuan, Ta-hsiang-ling, 2700 m, H.Smith 2083 (UPS); Nanchuan-hsien, c. 200–2200 m, W.P.Fang 974 (E); 1157 (E, K); Omei [Emei] Shan, E.H.Wilson 5303, 5304 (BM, K); Yunnan, Yangbi Xian, W side of Diancang Shan, 3300–3600 m, B.Bartholomew et al. 423 (E). TIBET: Pome [Bomi], Tongkah, F.Ludlow, G.Sherriff & R.H.J.Williams 13770 (BM). Extremely robust forms of var. chinensis are very different from the type variety but intermediates are known. 24. Luzula alpinopilosa (Chaix) Breistr., Bull. Soc. Sci. Isére 61: 609 (1947) Juncus alpinopilosus Chaix, in D.Villars, Hist. Pl. Dauphiné 1: 318 (1786). T: Haller 1326 [A. von Haller, Hist. Stirp. Helv. 2: no. 1326 (1768)]; syn: P-Haller; France, Chaliol-le-Viel, D.Chaix; syn: G, n.v. Luzula variabilis Buchenau, Bot. Jahrb. Syst. 7: 171 (1885), [‘species collectiva’], nom. illeg. Perennials 10–65 cm tall, subglabrous, loosely caespitose; rhizome branched, ascending; stolons often present, short. Basal leaves few, intermediate between cataphylls and cauline leaves; cauline leaves 3–5, 1.5–9.0 mm wide. All leaves with papillose-serrulate margins; apex acute. Lower bract herbaceous to subherbaceous, shorter than or rarely ±equalling inflorescence. Inflorescence a loose to subdense suberect or ±nodding panicle to 7 × 6 cm; flowers borne singly and in 2–3 (–7)-flowered clusters. Bracteoles ovate-lanceolate, 1.5–2.0 mm long, brownish, ciliate. Tepals ±equal, glabrous, lanceolate, with outer ones acuminate, 1.8–3.3 mm long, blackish brown to dark castaneous-brown; margins indistinct or very narrow. Stamens 6; anthers 0.6–1.2 mm long, 3–9 times longer than filaments; filaments 0.1–0.4 mm long; style 0.6–1.2 mm long; stigmas 1–2 mm long. Capsule ovoid to oblongovoid, subobtuse to ±acute, shortly mucronate, usually ±exceeding perianth, dark castaneousbrown, smooth; capsule segments usually 2.1–2.4 × 1.2–1.5 mm. Seeds pale brown, oblongovoid, 1.1–1.3 mm long, 0.6–0.7 (–0.9) mm wide; apex distinct, pale, to 0.1–0.2 mm long; basal appendage minute, as a paler patch to 0.1 mm long; base fibrillate. Mountains of C and S Europe. Very variable in size, leaf width, tepal length and anther/filament length ratio. Three subspecies are recognized but numerous intermediates exist. Subalpine and alpine grasslands, rocky slopes, on gravel along streams, confined to silicic substrates. 1 Filaments 0.1–0.2 (–0.3) mm long; anthers 4–9 times longer than filaments 24b. subsp. obscura 1: Filaments 0.3–0.4 mm long; anthers usually 2–4 times longer than filaments 2 Cauline leaves 4–7 cm long, 1.5–5.5 mm wide; anthers usually 0.9–1.2 mm long; lower bract substantially shorter than inflorescence 24a. subsp. alpinopilosa 2: Cauline leaves 7–13 cm long, 4–9 mm wide; anthers usually 0.7–0.8 mm long; lower bract ±equalling inflorescence or shorter 24c. subsp. deflexa 24a. Luzula alpinopilosa (Chaix) Breistr. subsp. alpinopilosa Juncus spadiceus All., Fl. Pedem. 2: 216 (1785), nom. illeg., non Vill. (1779); Luzula spadicea [All.] DC., Fl. Franç., 3rd edn, 3: 159 (1805); Luzula spadicea var. allionii E.Mey., Linnaea 22: 401 (1849), nom. inval.; Luzula spadicea var. barbata Neilr., Nachtr. Fl. Wien 100 (1851); Luzula variabilis Buchenau, Bot. Jahrb. Syst. 7: 171 (1885), [doubtfully valid: ‘species collectiva’], nom. illeg.; Juncoides spadicea Kuntze, Revis. Gen. Pl. 2: 724 (1891). T: Haller 1326; syn: P-Haller; Scheuchzer It. Alp. 6: 458; syn: n.v. [authentic material of Scheuchzer in W] Luzula spadicea var. laxiflora Desv., J. Bot. (Desvaux) 1: 146 (1809); Luzula spadicea f. laxiflora (Desv.) I.Grint., Fl. Rep. Soc. Roman. 11: 594 (1966). T: 'in pratis humidis Alpium Europae' [otherwise not indicated]. Luzula spadicea β [unranked] elatior Gaudin, Fl. Helv. 2: 569 (1828). T: [Switzerland] in valle Champé, J.Gay; syn: n.v. 53
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AUSTRALIAN BIOLOGICAL RESOURCES STU
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CONTENTS Contributors iv Species Pl
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Species Plantarum Steering Committe
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PREFACE This volume is the first of
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