Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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SPECIES PLANTARUM — FLORA OF THE WORLD (2002)
long; stigmas 1.0–1.2 mm long. Capsule subglobular, ±obtuse, mucronate, castaneous-brown;
capsule segments 1.7–2.0 mm long, including mucro 0.1–0.2 mm long and 1.4–1.5 mm wide.
Seeds subglobular, pale castaneous-brown, smooth, shining, 0.8–0.9 (–1.0) mm long, 0.7–0.8 mm
wide; apex pale; basal appendage indistinct, to 0.1 mm long, very sparsely fibrillate. 2n=12,
fide N.Galland, Trav. Inst. Sci. Univ. Mohammed V, Sér. Bot., 35 (22): 106–121 (1988).
Endemic to the Great Atlas and Anti-Atlas in Morocco. 20: MOR. Amongst shady mountain
rocks in open oak woodlands. Map 31.
20. MOROCCO: Great Atlas, Oukaimeden, 2500–2600 m, P.H.Davis 67595 (E); Reraya, Sidi-Chamarouch,
2350–2500 m, 1923, R.V.Litardiére (MPU); Great Atlas, Reraya Valley, Sidi-Chamarouch, 2400–2500 m,
21 Jul 1924, R.Maire (P); Great Atlas, Gourdouze, 1800 m, Apr 1928, J.Braun-Blanquet & I.Braun-Blanquet
(MPU).
A species in many respects intermediate between sections Anthelaea and Diprophyllatae.
The seed characters clearly point to Luzula (Marlenia) elegans Lowe.
Sect. 3. NODULOSAE
Luzula sect. Nodulosae Chrtek & Křísa, Preslia 46: 211 (1974)
Luzula subsect. Nodulosae (Chrtek & Křísa) Novikov, Byull. Moskovsk. Obshch. Isp. Prir., Otd. Biol. 95(6):
66 (1990). T: Luzula nodulosa E.Mey.
Perennials with thick nodulose horizontal rhizomes. Leaf tips acuminate, subulate. Lower
bract ±membranous to subherbaceous. Inflorescence umbelloid or anthelate, composed of
capituliform dense large clusters, on ±suberect peduncles. Seeds broadly ovoid, large,
1.7–1.9 × 1.3–1.4 mm, with apical and basal patches of looser outer seed-coat; appendages
absent; base distinctly fibrillate.
A monotypic section confined to the Mediterranean region, disjunctly in NW Africa and
from Greece to W Anatolia.
12. Luzula nodulosa [Bory & Chaub.] E.Mey., Linnaea 22: 410 (1849), nom. cons.
propos.
Juncus nodulosus Bory & Chaub., Exp. Sci. Morée Bot. 3: 105, t. 11, f. 1 (1832), nom. illeg., non Wahlenb.;
Juncus graecus Chaub. & Bory, Nouv. Fl. Pélop. 23, Plate 12, fig. 1 (1838); Luzula graeca (Chaub. & Bory)
Kunth, Enum. Pl. 3: 310 (1841); Juncoides nodulosa (E.Mey.) Kuntze, Revis. Gen. Pl. 2: 724 (1891); Luzula
nodulosa var. graeca (Chaub. & Bory) R.Maire, Bull. Soc. Hist. Nat. Afrique N. 22: 319 (1931). T: [Greece,
Pylos, Arcadia] L.A.Chaubard; holo: P.
Luzula nodulosa var. mauretanica R.Maire & Trab., in R.C.J.E.Maire, Bull. Soc. Hist. Nat. Afrique N. 22: 319
(1931). T: Algeria, Monts Tlemcen, entre Terni & Sebdou, L.Trabut; holo: P.
Illustration: F.G.P.Buchenau, in H.G.A.Engler, Pflanzenr. (iv.36) 25: 65, fig. 46a, 66, fig. 46b (1906).
Perennials c. (15–) 30–55 (–70) cm tall, loosely caespitose or one-stemmed; rhizome thick,
c. 3–5 mm in diam., creeping to ascending, often with fibrous cover, nodulose, with internodes
usually 0.5–2.0 cm long; stolons short, ascending. Lowermost leaves cataphyllous, c. 1–3 cm
long, brownish; basal leaves flat, 7–16 (–20) cm long, 3.5–5.0 (–7.5) mm wide; cauline
leaves 3–4 (–5), (1.5–) 2.5–5.0 (–15) cm long; margins densely ciliate, later sometimes
glabrescent, densely papillose-serrulate; apex subulate. Lower bract brownish-membranous
to subherbaceous, c. 1.0–3.5 (–5.0) cm long, shorter than inflorescence. Inflorescence of
(1–) 2–5 (–15) large hemispherical to cup-shapedclusters; clusters (2–) 3–5 (–11)-flowered,
0.7–1.0 cm long, c. 1.0–1.5 cm wide; peduncles suberect, c. 2–10 cm long, smooth;
secondary branches absent or frequent, distally inserted, short. Bracteoles scarious, often
brownish at base, ovate, acuminate, c. 3.0–3.5 mm long; midrib distinct. Tepals ±equal,
ovate-lanceolate or broadly lanceolate, c. 5.0–6.5 mm long, acute to acuminate; central band
deep castaneous-brown; margins distinct, broad, membranous. Stamens 6; anthers 1.7–3.5 mm
long, 2–10 times as long as filaments; filaments 0.3–0.9 mm long; style 1.8–2.7 mm long;
stigmas c. 2.5–4.0 mm long. Capsule ovoid to broadly ovoid, acuminate, shorter than
perianth, dark castaneous to pale castaneous-brown; capsule segments 3.3–4.0 × 2.3–2.8 mm