54 SPECIES PLANTARUM — FLORA OF THE WORLD (2002) <strong>Luzula</strong> spadicea var. candollei E.Mey., Linnaea 22: 401 (1849); <strong>Luzula</strong> spadicea subsp. candollei (E.Mey.) K.Richt., Pl. Eur. 1: 185 (1890); <strong>Luzula</strong> alpinopilosa subsp. candollei (E.Mey.) Rothm., Feddes Repert. Spec. Nov. Regni Veg. 67: 4 (1963). T: France, E Pyrenees, Val d’Eynes, 13 Sep 1839, R.J.Shuttleworth; syn: W; Switzerland, Geneve, Castan; syn: n.v.; Pyrenees, Meles, L.Marchand; syn: n.v. Illustrations: H.E.Hess, E.Landolt & R.Hirzel, Fl. Schweiz 1: 519 (1967), as L. spadicea; S.Pignatti, Fl. Ital. 3: 445 (1982). Plants (10–) 15–20 (–35) cm tall. Leaves of sterile shoots (when present) usually 5–10 (–20) cm long, 2–4 (–5) mm wide; cauline leaves usually 4–7 cm long, 1.5–3.5 (–5.5) mm wide. Lower bracts subherbaceous or pale brownish, 2–3 (–4) cm long, substantially shorter than inflorescence. Inflorescence usually 4–7 × 3–6 cm, suberect <strong>to</strong> slightly nodding, ±evenly branched, most flowers usually borne singly, but with 2–4-flowered clusters usually also present. Tepals 1.8–2.7 mm long. Anthers 0.9–1.2 mm long, usually 3–4 times longer than filaments; filaments 0.3–0.4 mm long; style (0.7–) 0.9–1.2 mm long; stigmas c. 1.5 mm long. 2n=12 [AL], fide H.Nordenskiöld, Hereditas 37: 330–331 (1951); Druskovic, Int. Organ. Pl. Biosyst. Newslett. 24: 13 (1995). Mountains of Central and southern Europe: the Alps, the N Apennines, the Pyrenees, the Vosges. 11: AUT, GER, SWI. 12: FRA, SPA. 13: ITA, YUG-SL. Rocky and s<strong>to</strong>ny subalpine and alpine slopes. Map 47. 11. AUSTRIA: Steiermark, Bösenstein, 1896 m, G.Strobl [Fl. Exs. Austro-Hung.] 1872 (PR, PRC, WU); Vorarlberg, Montafon, Hochmaderjoch, Gölles 462 (WU). SWITZERLAND: Tessin, St. Gotthard, Monte Prosa, 2400 m, A.Kneucker [Cyper. Junc. Exs.] 90 (PR, PRA); Valais, Zermatt, Riffelberg, F.O.Wolf [A.Kneucker, Cyper. Junc. Exs.] 90a (PR). 13. ITALY: N Apennines, Corno alle Scale, 21 Jul 1840, P.Savi (PR); Saint-Rhémy, 2400 m, A.Charpin, W.Greuter & Hainard [Exs. Genav.] 117 (W). Very variable but the two extremes (the broad-leaved, small-flowered form often recognized as subsp. candollei, and the typical narrow-leaved large flowered plants) often coexist within populations, and intermediates are not rare. 24b. <strong>Luzula</strong> alpinopilosa subsp. obscura S.E.Fröhner, Preslia 40: 426 (1968) <strong>Luzula</strong> obscura (S.E.Fröhner) Novikov, Byull. Moskovsk. Obshch. Isp. Prir., Otd. Biol. 95(6): 66 (1990). T: Slovakia, Vysoké Tatry, Zbojnická chata, 1900 m, 14 Aug 1965, S.Fröhner; holo: PR. <strong>Luzula</strong> carpatica Kitt. ex Kanitz, Linnaea 32: 327 (1863), an nom. validum?; <strong>Luzula</strong> spadicea A.b. [unranked] carpatica (Kitt. ex Kanitz) Asch. & Graebn., Syn. Mitteleur. Fl. 2(2): 513 (1904);. <strong>Luzula</strong> spadicea f. carpatica (Kitt. ex Kanitz) I.Grint., Fl. Rep. Soc. Roman. 11: 594 (1966). T: [Slovakia], 'in der kleinen Kohlbach', J.D.T.Mauksch; holo: n.v., probably BP. Illustration: T.Szynal & J.Mądalski, Atlas Fl. Polsk. 2(2): 151 (1931). Plants 10–30 cm tall. Cauline leaves 3–8 cm long, 3.5–<strong>6.</strong>5 mm wide. Lower bract usually herbaceous, usually 1.5–4.5 cm long, shorter than inflorescence. Inflorescence c. 4–5 × 3–5 cm, ±nodding <strong>to</strong> suberect; main branches long; secondary branches often abbreviated; flowers borne singly and usually in (2–) 3–5 (–7)-flowered loose clusters. Tepals (2.0–) 2.2–3.0 mm long. Anthers 0.9–1.2 mm long, usually 4–9 times longer than filaments; filaments 0.1–0.2 (–0.3) mm long; style 0.8–0.9 mm long; stigmas 1.2–2.0 mm long. 2n=12 [AL], fide J.Májovský & A.Murín, Karyotax. Prehl. Fl. Slov. 361 (1987). Fig. 12. C Europe, confined <strong>to</strong> the Carpathians from Slovakia and Poland <strong>to</strong> Romania. 11: CZE-SL, POL. 13: ROM. 14: UKR. Map 48. 11. SLOVAKIA: Nízké Tatry, Králova Hola, 1700 m, 20 Aug 1898, K.Tocl (PR); Vysoké Tatry, Mlynická dolina, 1700 m, 10 Jul 1925, K.H.Rechinger (W). 13. ROMANIA: Retezat [Mtns], Gura Apei, Zanoga Lake, 2 Jul 1970, J.Soják (PR); Muntii Fogarasului, Vistea, Sambatei Valley, 1200–2000 m, 30 Jun 1970, J.Soják (PR). 14. UKRAINE: the East Carpathians, Mt Goverla, 2030 m, 7 Jul 1948, E.Bradis & A.Zanja<strong>to</strong>va (MW); Zakarpatskaya area, Rachiv, Mt Turkul (1850 m) 16 km NE of Bogdan, 1740 m, 27 Jul 1996, Z.Kaplan 96/506 (PRA). The overall variation of subsp. obscura is greater than that described in the pro<strong>to</strong>logue. In particular, narrow-leaved and small-flowered plants occur in populations of subsp. obscura. Some plants from Bulgaria (the Stara Planina Mtns) are intermediate between subsp. obscura and subsp. deflexa.
<strong>JUNCACEAE</strong> (<strong>Luzula</strong>) Figure 12. <strong>Luzula</strong> alpinopilosa subsp. obscura. A, habit; B, inflorescence; C, terminal group of flowers; D, flower; E, capsule; F, seed; G, tepal; H, bracteole; I, stem T.S.; J, leaf T.S. (A, C–J, Piasecki, 4 Aug 1890, KRA; B, A.Rehman s.n., KRA). Scale bars: A, B = 5 cm; C = 5 mm; D–H = 2 mm; I, J = 1 mm. Drawn by T.Szynal. Reproduced with permission, from T.Szynal & J.Mądalski, Atlas Fl. Pol. 2/2: 151 (1931). 55
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