Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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74 SPECIES PLANTARUM — FLORA OF THE WORLD (2002) 1370 m, 29 Jan 1955, I.Skottsberg & C.J.F.Skottsberg 399 (S); Masafuera, Cuchara, above Correspondencia Camp, 1350 m, I.Skottsberg & C.J.F.Skottsberg 162 (S); Masafuera, Quebrada Guaton, valley toward Cordon Atravesado, 1230 m, A.Landero & L.Gaete 9108 (OS); Masafuera, 1200–1300 m, C.J.F.Skottsberg 28 (UPS). 36. Luzula alopecurus Desv., J. Bot. (Desvaux) 1: 159 (1808) Juncoides alopecurus (Desv.) Kuntze, Revis. Gen. Pl. 2: 734 (1891). T: ‘ad fretum magellanicum’; 'Magell.', P.Commerson; syn: P. Luzula villosa Wikstr., Kongl. Vetensk. Akad. Handl. 1823(2): 276 (1823). T: ‘Terra del Fuego et Falklands Islands’; syn: not located in S or UPS. Illustrations: F.G.P.Buchenau, in H.G.A.Engler, Pflanzenr. (iv.36) 25: 78, fig. 50 (1906); M.Barros, Darwiniana 10: 339, fig. 18 (1953). Map: H.Meusel, E.Jäger & E.Weinert, Vergl. Chorol. Zentraleur. Fl. 1: 88 (1965). Perennials usually 10–35 cm tall, densely hairy caespitose; base often purplish, robust; rhizome short; stolons absent. Stem rigidly erect, (0.6–) 0.8–1.2 mm in diam. below inflorescenceBasal leaves densely ciliate, numerous, 8–15 (–18) cm long, 4–5 (–6) mm wide; cauline leaves unevenly distributed, usually with one near stem base and 1 or 2 close to inflorescence, 6–10 cm long, often overtopping inflorescence; all leaves acute, with margins papillose-serrulate. Lower bract herbaceous, c. 2–5 cm long, usually overtopping inflorescence; upper bracts scarious-brownish, fimbriate-ciliate, up to 2 cm long. Inflorescence erect, densely congested, c. 1.5–2.5 cm long, of whitish villose appearance when younger, rarely with a small separate flower cluster at base. Bracteoles inserted c. 0.5–1.0 mm below flower; upper bracteoles narrowly lanceolate with a filiform tip, c. 3.0–3.5 mm long (often as long as flowers), densely fimbriate–ciliate. Tepals subentire, with outer ones densely and inner ones usually very sparsely long ciliate, narrowly lanceolate, acuminate with filiform tip, ±unequal (inner tepals 2.5–2.9 mm long, outer tepals 2.9–3.7 mm long), purplish castaneous brown to straw brown at base, with broad membranous border. Stamens 6; anthers 0.4–0.7 mm long; filaments 0.5–0.9 mm long, longer than anthers; style 0.1–0.2 mm long; stigmas c. 1.0 mm long. Capsule oblong-ovoid, ±trigonous above, ±acute, castaneous brown, shorter than perianth; capsule segments 1.8–2.0 × 1.3–1.4 mm. Seeds ellipsoidal, with distinct yellowish seed-coat and yellowish apex, dark castaneous, 0.9–1.0 mm long, 0.5–0.6 mm wide; appendage yellowish, 0.1–0.2 mm, fibrillate. 2n=24, fide D.M.Moore, Fl. Tierra del Fuego 325 (1993). Endemic to southernmost Patagonia, Tierra del Fuego and the Falkland Islands. 85: AGS, CLS. 90: FAL. Grasslands and rocky slopes. Map 66. 85. ARGENTINA SOUTH: Santa Cruz, C.Burmeister 173 (SI); Tierra del Fuego, Ushuaia, Río Piper, R.Santesson 411 (S); Isla Soledad, Seal Rookery, E.A.Ulibarri, M.Dimitri & E.Orfila 1039 (SI). CHILE SOUTH: Magallanes, Punta Arenas, Loretto, 9 Dec 1903, Scott Elliott (BM); Tierra del Fuego, Porvenir, R.Santesson 1565 (S); Tierra del Fuego, Sierra Carmen Silva, N side of Río Chico valley, D.M.Moore & R.N.Goodall 225 (H, RNG, UPS). 90. FALKLAND IS.: West Falkland Is., Roy Cove Ck, Vallentin 84 (K); Port Stanley, C.J.F.Skottsberg 90 (BM); 4 Jan 1908, C.J.F.Skottsberg 28 (S); East Falklands, Kidney Is., Sladen 5/49 (BM). 37. Luzula brachyphylla Phil., Linnaea 33: 268 (1865) Luzula racemosa f. brachyphylla (Phil.) Fuentes, Bol. Mus. Nac. Hist. Nat. 90: 137 (1917). T: Chile, Chiloë Is., Chacao, Fonck 44; holo: SGO. Luzula patagonica Speg., Rev. Fac. Agron. Univ. Nac. La Plata 3: 577 (1897); Juncoides patagonica (Speg.) Kuntze, Revis. Gen. Pl. 2: 734 (1891). T: Argentina, Santa Cruz, Lago Argentino, Arroyo Leona, C.Spegazzini; syn: S. Luzula chilensis f. frequentior Barros, Darwiniana 10: 321 (1953). T: Argentina, Chubut, valle Lonquíñeu, Los Menucos, 8 Feb 1945, A.Castellanos; lecto: LIL114298, designated here. Luzula correae Barros, in M.N.Correa, Fl. Patagon. 2: 118 (1969). T: Argentina, Santa Cruz, Calafate, M.N.Correa 2962, L.Mendonza & C.Movia; holo: BAB, photo PRA. Perennials 15–30 (–55) cmtall, loosely caespitose, sparsely hairy; rhizome short or ascending; stolons absent. Basal leaves 4–8 (–17) cm long, 3–4 mm wide; cauline leaves 1 or 2, usually 3–7 cm long; all leaves ±acute, with papillose-serrulate margins. Bracts erect to erecto-
JUNCACEAE (Luzula) patent, 1.5–2.5 (–4.0) cm long, equalling or overtopping their partial inflorescence, with lower bracts herbaceous and upper ones scarious. Inflorescence erect, rarely slightly nodding, interrupted, 2.5–6 cm long, of 3–6 (–10) clusters, with lower clusters pedunculate and upper ones subsessile; all clusters elongated, narrow (c. 4 mm in diam.), cylindrical composed of subclusters, many-flowered. Bracteoles c. 1 mm long, ovate, sparsely ciliate, scarious. Tepals ±lanceolate, subequal, glabrous; outer tepals acuminate, subaristate; inner tepals ±mucronate, 1.7–2.0 (–2.3) mm long; base castaneous; margin paler or membranous. Stamens 3 or 4–6; anthers 0.5–0.8 mm long; filaments 0.3–0.4 mm long; style 0.1–0.2 mm long; stigmas c. 1 mm long. Capsule ±ovoid, ±acute, deep brown; capsule segments 1.6–1.8 × 1.0–1.1 mm long. Seeds dark, ellipsoidal, 0.9–1.0 (–1.1) mm long, 0.6–0.7 mm wide; seedcoat not distinct; apex not distinct; appendage c. 0.05 mm long, fibrillate. Fig. 17. Southern Chile and Southern Argentina. 85: AGS, CLS. In grasslands, usually at lower altitudes. Map 67. 85. ARGENTINA SOUTH: Tierra del Fuego, Río Azopardo, P.Dusén 584 (UPS); Laguna Patos near Río Carbon, 4 Jan 1909, C.J.F.Skottsberg 943 (S, UPS); Chubut, Trevelin, T.Meyer 9277 (LIL); Chubut, José de San Martín, 530 m, 22 Jan 1902, Hözberg (S); Neuquén, Ñorquín, Copahué, 2000 m, C.A.O’Donnell 2097 (LIL); Neuquén, Pino Hachado, A.Burkart 9651 (SI). CHILE SOUTH: Magallanes, Cerro Paine, S.F.Anilot 6094 (SGO); Aysén, Laguna San Rafael, 3–4 m, C.Muñoz 6147 (SGO); Los Lagos, Mechuque, F.W.Pennel 12622 (SGO); Los Lagos, San Rafael, 1921, C.M.Hicken (SI 10974). Most plants from Chile and from the southern part of the range are triandrous, with occasional occurrence of flowers with 4–6 stamens. Plants from Santa Cruz, Chubut and Neuquén, Argentina, are predominantly hexandrous (flowers with 3–5 stamens sometimes found in the same inflorescence as hexandrous ones). Differences in stamen number is the only significant variation found within the species (the names L. patagonica and L. correae refer to the predominantly hexandrous form). 38. Luzula chilensis Nees & Meyen ex Kunth, Enum. Pl. 3: 312 (1841) Juncoides chilensis (Kunth) Kuntze, Revis. Gen. Pl. 2: 734 (1891). T: Chile, ‘ad Maipú fluv., alt. 10000 ped.’, 1831, Meyen; syn: B, destroyed, see F.G.P.Buchenau, Abh. Naturwiss. Vereine Bremen 6: 418, Plate iv, fig. 1, a–d (1879) [et Junc. S.-Amer., 1879]; Chile, Bíobío, Antuco, E.F.Poeppig [Coll. Pl. Chil. III] 105; syn: BM, P, S. Luzula rigida Phil., Linnaea 33: 267 (1865); Luzula chilensis f. rigida (Phil.) Fuentes, Bol. Mus. Nac. Hist. Nat. 90: 138 (1917). T: Chile, Colchagua, ‘in montibus subalpinis Talcaregué, Feb 1831, [J.Gay] 284; holo: SGO. Luzula chilensis f. longifolia Barros, Darwiniana 10: 322 (1953). T: Argentina, Neuquén, Río Alumine, Otto Asp. 98; syn: BAF; Chubut, Río Percey, 6 Feb 1945, A.Castellanos; syn: LIL114302. Perennials, usually 12–25 (–55) cm tall, caepitose, sparsely to ±densely hairy; rhizome short; stolons absent. Basal leaves (5–) 6–8 (–12) cm long, 2.5–3.5 (–6) mm wide; cauline leaves 2, usually short, 1.5–3.0 cm long; all leaves acute, with margins distinctly papillose-serrulate. Lower bract membranous-brownish, rarely herbaceous, usually 1.5–2.0 cm long, ciliate, ±equalling inflorescence. Inflorescence usually congested as a single slightly nodding or ±suberect head c. 2 × 1.5 cm, of 5–6 clusters (usually 3–6-flowered); less often clusters distinct, ±interrupted, inflorescence about 5 cm long. Bracteoles scarious with straw-brown base, ovate, acuminate, fimbriate-ciliate, 1.5–2.0 mm long. Tepals ±equal, glabrous, lanceolate, c. 2.7–3.3 mm long, acuminate-aristate, pale castaneous brown below and on midrib, otherwise membranous to pale straw-brown. Stamens 6; anthers usually 0.6–0.9 mm long; filaments 0.4–0.5 mm long; style 0.3–0.4 mm long; stigmas 1.2–1.6 mm long. Capsule ovoid to subglobular, ±obtuse, castaneous brown, shining, equalling or shorter than perianth; capsule segments 2.4–3.0 × 1.5–2.1 mm. Seeds paler castaneous brown; seed-coat distinct, forming a yellowish apex, 1.2–1.5 mm long, (0.9–) 1.0–1.2 mm wide; appendage (0.1–) 0.2 mm long, fibrillate. Fig. 17. Most common in S and C Chile and S and W Argentina, reaching Valparaiso (district of Aconcagua) in C Chile. 85: AGS, AGW, CLC, CLS. Distribution imperfectly known. Wet mountain slopes and grasslands, usually below 2000 m, rarely above 3000 m. Map 68. 75
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AUSTRALIAN BIOLOGICAL RESOURCES STU
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CONTENTS Contributors iv Species Pl
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Species Plantarum Steering Committe
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PREFACE This volume is the first of
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