30 SPECIES PLANTARUM — FLORA OF THE WORLD (2002) 9b. <strong>Luzula</strong> luzuloides subsp. rubella (Hoppe ex Mert. & W.D.J.Koch) Holub, Folia Geobot. Phy<strong>to</strong>tax. 18: 205 (1983) <strong>Luzula</strong> albida var. versicolor Bluff & Fingerh., Comp. Fl. German. 1: 443 (1825); <strong>Luzula</strong> albida var. rubella Hoppe ex Mert. & W.D.J.Koch, in Deutschl. Fl.(Röhling) 2: 599 (1826); <strong>Luzula</strong> nemorosa var. rubella (Hoppe ex Mert. & W.D.J.Koch) Garcke, Fl. N. Mitt.-Deutschland, 2nd edn, 340 (1851); <strong>Luzula</strong> angustifolia var. rubella (Hoppe ex Mert. & W.D.J.Koch) Garcke, Fl. N. Mitt.-Deutschland, 3rd edn, 348 (1853); <strong>Luzula</strong> nemorosa f. rubella (Hoppe ex Mert. & W.D.J.Koch) Racib., Consp. Juncac. Polon. 25 [p. separ.] (1888); <strong>Luzula</strong> angustifolia subsp. rubella (Hoppe ex Mert. & W.D.J.Koch) K.Richt., Pl. Eur. 1: 184 (1890); <strong>Luzula</strong> albida subvar. rubella (Hoppe ex Mert. & W.D.J.Koch) Rouy, Fl. France 13: 261 (1912). T: Austria, Heiligenblut, D.H.Hoppe [Herb. Viv. Pl. Gram.] 68; syn: LI, BRNM etc. (further authentic material: K, LD, PR). <strong>Luzula</strong> albida var. cuprina Rochel, Pl. Banat. Rar. 3: 27 (1828), nom. inval.; <strong>Luzula</strong> nemorosa A.II.b. [unranked] cuprina [Rochel ex] Asch. & Graebn., Syn. Mitteleur. Fl. 2(2): 503 (1904); <strong>Luzula</strong> luzuloides subsp. cuprina [Rochel ex] Chrtek & Křísa, Preslia 46: 212 (1974). T: Slovakia, 'ad pedes Kralowahora', A.Rochel; authentic material: W, PR [as Juncus albidus var. cuprinus]. <strong>Luzula</strong> nemorosa var. colorata E.Mey., Linnaea 22: 395 (1849). T: in Alpibus. D.H.Hoppe; syn: LI; in Carpathis, Baumgarten; syn: n.v. <strong>Luzula</strong> alpigena Schur, Enum. Pl. Transsilv. 681 (1866). T: n.v. [numerous syntypes from Romanian South Carpathians, most probably in LW]. <strong>Luzula</strong> albida var. erythranthema Wallr., Sched. Crit. 1: 150. (1822); <strong>Luzula</strong> nemorosa var. erytranthema (Wallr.) Hayek, Fl. Stir. Exs., no. 409 (c. 1930) [printed exsiccatum label]; <strong>Luzula</strong> luzuloides var. erythranthema (Wallr.) I.Grint., Fl. Rep. Soc. Roman. 11: 592 (1966). T: Austria, Heiligenblut, D.H.Hoppe; syn: PR [specimen annotated by C.F.W.Wallroth]; Heiligenblut, D.H.Hoppe [Herb. Viv. Pl. Gram.] 68; syn: LI, BRNM. <strong>Luzula</strong> fuscata Schur, Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 10: 200 (1859). T: [Romania], 'Am Büdöskegel', F.Schur; holo, n.v. [doubtfully valid]. <strong>Luzula</strong> nemorosa [unranked] fuliginosa Asch., Fl. Brandenburg 1: 740 (1864); <strong>Luzula</strong> albida var. fuliginosa (Asch.) Beck, Fl. Nieder-Österreich 1: 159 (1890). T: [Czech Republic, the Krkonoše, Labský vodopád], am Elbfall des Riesengebirges, Jul 1862, C.Lucas; holo: n.v. <strong>Luzula</strong> nemorosa f. rosea Kožuharov, Izv. Bot. Inst. (Sofia) 11: 127 (1963), nom. inval. <strong>Luzula</strong> nemorosa f. grandiflora Kožuharov, Izv. Bot. Inst. (Sofia) 11: 127 (1963), nom. inval., & in D.Jordanov, Fl. N. R. B'lgar. 2: 401 (1964), nom. inval. T: Bulgaria, Mt Balkan centralis [collec<strong>to</strong>r not given]; holo: n.v. <strong>Luzula</strong> nemorosa f. aggregata Kožuharov, Izv. Bot. Inst. (Sofia) 11: 127 (1963). T: Bulgaria, Vi<strong>to</strong>sha, A.Yavashev; holo: SOM. <strong>Luzula</strong> luzuloides subsp. tenacissima Vivant, Bull. Soc. Bot. France 126: 259 (1979). T: France, W Pyrenees, Haute-Soule, Pic Lackhoura, 1700–1800 m, 5 Sep 1973, J.Vivant s.n.; holo: P; iso: JACA. Illustration: T.Szynal & J.Mądalski, Atlas Fl. Polsk. 2(2): 152 (1931). Plants loosely <strong>to</strong> ±densely caespi<strong>to</strong>se; s<strong>to</strong>lons short, ascending. Bracteoles usually pale castaneous or coppery-coloured. Tepals usually coppery-coloured, brownish-purplish or pale castaneous with reddish tinge, usually 3.0–3.5 (–4.2) mm. 2n=12, fide C.Garcia-Herran, in litt. (material from Polish Tatra Mtns and Bieszczady, Poland, vouchers in NEU). Fig. 7. Mountains of C and S Europe, the Pyrenees, introduced <strong>to</strong> Great Britain. 10: grb. 11: AUT, CZE, GER, POL, SWI. 12: FRA, SPA. 13: ALB, BUL, GRC, ITA, ROM, YUC. 14: UKR. Mountain <strong>to</strong> subalpine tall-grass meadows, open mountain woodlands; in mountain regions where it is common, it is also found in valley situations. Map 2<strong>6.</strong> 11. AUSTRIA: Oetztaler Alpen, Landeck, Venetberg, 2300 m, Aug 1968, A.Polatschek (W); Steiermark, Pretulalpe, Fischbacher Alm, 16 Jul 1911, K.Ronniger (W). CZECH REPUBLIC: Hrubý Jeseník, Petrovy kameny, 27 Jul 1889, V.Spitzner (PR 49587). POLAND: Babia Góra, 1480 m, Walas [Pl. Polon. Exs.] 81 (K). 13. BULGARIA: Vi<strong>to</strong>ša, J.Velenovský (PR 49584). CRNA GORA: Sekirica Planina, Jul 1903, J.Rohlena (PR). ITALY: South Tyrol, Dolomiti, Villnoss, Zamseralpe, 1912, K.Ronniger (W). F.Y.R. MACEDONIA: Skopje, Šar Planina, Ljuboten, 12 Jul 1959, J.D.A.Stain<strong>to</strong>n 7988 (K). Variable in leaf width, size of inflorescence and perianth colour. Intermediates between subspecies are not rare, e.g., 11. POLAND: Nowy Targ, Pieniny, S<strong>to</strong>larzówka, 620–640 m, H.Piękoś [Fl. Polon. Exs.] 389 (KRA, KRAM, W). Maybe better treated as a variety.
<strong>JUNCACEAE</strong> (<strong>Luzula</strong>) Figure 7. <strong>Luzula</strong> luzuloides subsp. rubella. A, habit; B, terminal part of inflorescence; C, H, flowers; D, capsule; E, seed; F, G, tepals; I, stem T.S.; J, leaf T.S. (A–C, F–I, A.Wołoszczak, Aug 1887, KRA; D, E, J.M ądalski s.n., 10 Aug 1927, KRA). Scale bars: A = 5 cm; B = 5 mm; C–H = 2 mm; I, J = 1 mm. Drawn by T.Szynal. Reproduced with permission, from T.Szynal & J.Mądalski, Atlas Fl. Pol. 2/2: 152 (1931). 31
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