Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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78 SPECIES PLANTARUM — FLORA OF THE WORLD (2002) Luzula macusaniensis Steud. ex Buchenau, Abh. Naturwiss. Vereine Bremen 4: 131 (1874); Juncoides macusaniensis (Buchenau) Kuntze, Revis. Gen. Pl. 2: 734 (1891).. T: Peru, Puno, Macusani, 1854, W.Lechler 1839; lecto: P, fide H.Balslev, Fl. Neotrop. Monogr. 68: 467 (1996) (as ‘holotype’); isolecto: BR, G, GOET, K, P, S, WRSL. Illustrations: N.A.Desvaux, J. Bot. (Desvaux) 1: 162, tab. 6, fig. 3 (1808); H.Balslev, Fl. Neotrop. Monogr. 68: 66, fig. 34, C, D (1996). Perennials (5–) 10–35 (–40) cm tall, caespitose, hairy; rhizome short, erect or ascending; stolons absent. Stem thin, nodding in upper part. Basal leaves canaliculate or flat, 5–18 cm long, typically 2.0–2.5 (–4.0) mm wide; cauline leaves (0–) 1 or 2, usually remote, narrow, usually 3.5–7 cm long; all leaves acute, with margins papillose-serrulate. Lower bract usually herbaceous, very narrow, up to 4–5 cm long, shorter than inflorescence; bracts supporting flower clusters linear-lanceolate, pale brownish-membranous, ciliate, up to 1 cm long, usually shorter than clusters. Inflorescence nodding, spike-like, (1.0–) 1.5–5 × 1.0–1.5 cm, of 3–8 elongated clusters, ±interrupted (rhachis visible), rarely compactly congested in high mountain forms (then only 1–1.5 cm long). Bracteoles ±membranous, 1.2–1.5 (–2.0) mm long, ovate, acuminate, ciliate. Tepals ±unequal, ±glabrous (rarely with few teeth or hairs), dark castaneous brown with paler apex and margins; outer tepals usually 2.0–2.4 (–3.0) mm long, lanceolate, acuminate-aristate; inner tepals 1.7–2.1 (–2.4) mm long. Stamens 3, rarely 4–5; anthers 0.3–0.5 mm long; filaments 0.3–0.6 mm long; style 0.1–0.2 mm long; stigmas 0.6–0.9 mm long. Capsules oblong-ovoid, usually ±obtuse, shorter than perianth, paler brown; capsule segments usually 1.5–1.7 (–1.8) × 0.9–1.1 mm. Seeds pale brown, shining, with darker apex, (0.7–) 0.9–1.0 mm long, c. 0.6 mm wide; appendage distinct, yellowish, 0.1–0.2 mm long, sparsely short fibrillate. 2n=24, fide J.H.Beaman, Amer. J. Bot. 49(1): 41–50 (1962), n.v. Fig. 18. Central (C Mexico and Guatemala) and South America, from Venezuela to Argentina and Chile. 79: MXC, MXG, MXS. 80: GUA, SMX. 82: VEN. 83: BOL, CLM, ECU, PER. 85: AGS, AGW, CLN, CLS. Sunny and rocky sites in open woodlands and páramo, usually between 3000 and 4000 m. Map 70. 79. MEXICO: Puebla, N side of Popocatépetl, J.H.Beaman 1742 (DUKE, GH, TEX, US). 80. GUATEMALA: San Marcos, Totonicapan, Tecum Uman ridge, 3340 m, J.H.Beaman 4180 (DUKE, NY, UC). 82. VENEZUELA: Mérida, Libertador, Sierra Nevada, Loma Redonda, 4045 m, L.J.Dorr & L.C.Barnett 5213 (AAU, NY). 83. BOLIVIA: La Paz, Inquisivi, source of Rio Ocsalla, 3800 m, M.Lewis 40912 (AAU, LPB, MO). COLOMBIA: Páramo Choachi, 3000 m, A.Lindig 1404 (P, S). ECUADOR: Cotopaxi, 4 km S of Cotopaxi railway station, 3500 m, H.Balslev 1183 (AAU, NY). PERU: Huaráz, Huascarán, Quebrada Churup, 3890–4260 m, D.N.Smith, Valencia & A.Gonzales 9629 (AAU, MO). 85. ARGENTINA NORTHWEST: Jujuy, Santa Ana, A.Burkart & N.S.Troncoso (SI11763); Tucumán, Ciénaga Grande, Cumbres Calchaquies, 3800 m, S.Halloy s.n. (LIL567036). ARGENTINA SOUTH: Neuquén, Los Lagos, San Martín de los Andes, A.E.Rasp 6 (SI) An extremely variable species in most features of general habit. In various regions plants have features blurring the otherwise distinct limits of the species. In Mexico, flowers are close to the upper size limit, in Ecuador, an intermediate between L. racemosa and L. chilensis was found (Sayausí, G.Harling 1424, S). Plants in Argentina have a tendency towards longer peduncles of lower clusters, a feature found in related species. The plasticity of vegetative traits, depending often on available moisture and altitude, is also enormous. The whole complex is in a need of a detailed cultivation and karyological study. 41. Luzula peruviana Desv., J. Bot. (Desvaux) 1: 160 (1808) Juncoides peruviana (Desv.) Kuntze, Revis. Gen. Pl. 2: 725 (1891). T: ‘in Terra Patagonala et montibus frigidetis Peruviae Pumbo, Antisana et Soluca.’; America calidiore, sine coll.; lecto: P, fide H.Balslev, Fl. Neotrop. Monogr. 68: 67 (1996) 1996: 67 [as holotype, left hand specimen typical]; Ecuador, Antisana, A.Humboldt; para: B-WILLD. Luzula boliviensis Buchenau, Abh. Naturwiss. Vereine Bremen 4: 128 (1874); Juncoides boliviensis (Buchenau) Kuntze, Revis. Gen. Pl. 2: 724 (1891). T: Bolivia, La Paz, Larecaja, Sorata, between Pongo and Anilaya, 3800 m, G.Mandon 1454; lecto: GOET, fide H.Balslev, Fl. Neotrop. Monogr. 68: 67 (1996); isolecto: BR, G, NY, P, S, UPS, US [p.p.; a few plants belong to L. racemosa or to intermediates]. Luzula peruviana var. longispica Barros, Darwiniana 10: 346 (1953). T: Argentina, Jujuy, Volcán, Loma del Tambo, 2500–3000 m, 22 Feb 1924, Schreiter 2909; holo: LIL.
JUNCACEAE (Luzula) Illustration: H.Balslev in Fl. Ecuador 11: 17, fig. 5 (1979). Perennials up to 60 cm tall, caespitose, hairy; rhizome vertical or ascending; stolons absent. Stem thick, rigidly erect, with only the uppermost part nodding. Basal leaves numerous, often squarrose, rigid, flat or canaliculate, usually 5–10 cm long and 2–4 mm wide; cauline leaves 2–4, densely hairy, to 13 cm long, evenly distributed on stem, often overlapping; all leaves acute, indistinctly papillose-serrulate. Lower bract herbaceous, densely hairy, up to 5 cm long, overtopping inflorescence; middle bracts supporting flower clusters purplish-brownish membranous, lanceolate, aristate, densely ciliate, c. 0.8–1.5 (–2.0) cm long, often exceeding the partial inflorescence. Inflorescence erect to slightly nodding, compact (rhachis usually not visible), usually 2.5–5 × c. 1.5 cm, often lobulate, rarely interrupted at base, dark castaneous brown, composed of c. 3–8 dense clusters. Bracteoles castaneous-membranous, ovatelanceolate, aristate, 2.0–2.5 mm, long ciliate. Tepals subequal or irregularly unequal, lanceolate, acuminate and aristate, usually ciliate, dark castaneous brown; upper part membranous with prominent midrib; longest tepals in flower 3.0–4.4 mm long. Stamens 3; anthers 0.3–0.4 mm long; filaments 0.6–0.9 mm long; style 0.1–0.2 mm long; stigma 0.5–1.1 mm long. Capsule oblong-ovoid, subacute to obtuse, paler brown; capsule segments 1.8–2.2 × 1.3–1.4 mm. Seeds paler brown, with darker apex, ellipsoidal, 1.1–1.4 mm long, c. 0.7 mm wide; appendage scarcely visible, c. 0.1 mm long, densely long fibrillate (seeds difficult to release from capsule). Centred to Ecuador and Peru, rare in Colombia, Bolivia and northernmost Argentina, with an isolated station in C Mexico; further exploration needed. 79: MXC. 83: BOL, CLM, ECU, PER. 85: AGW. In páramos, grasslands, rocky slopes, usually between 3800 and 4500 m. Map 71. 79. MEXICO CENTRAL: Tlaxcala, Malinche, 4400–4450 m, J.H.Beaman 2223 (UPS). 83. BOLIVIA: La Paz, Murillo, Laguna Zongo, 4740 m, J.Brandbyge 582 (AAU). ECUADOR: Chimborazo, Mt Chimborazo, 4550 m, E.Asplund 7938 (S, UPS); Pichincha, Páramo de Guamani, 3800–4100 m, H.Balslev 1618 (AAU); Imbabura, Volcán Cotocachi, 3900–4100 m, S.Laegaard 54498 (AAU). PERU: Puno, Araranca, 4100–4300 m, F.W.Pennell 13456 (S); Huaron, 4500 m, J.F.Macbride & F.Featherstone 1132 (S); Ancash, Carhuaz, Quebrada Ulta, 4400–4600 m, D.N.Smith 11338 (AAU, MO). 85. ARGENTINA NORTHWEST: Salta, Santa Victoria, Cochita, Ruta 5, foot of cerro Campanario, 30-35 km W of Santa Victoria, 4100–4200 m, L.J.Novara 6683 (MCNS); Tucumán, Cumbres Calchaquíes, Callejones, 18 Feb 1915, L.Castillón (LIL 50636). Variable, and in many features similar to L. racemosa. Main diagnostic characters are its ciliate longer tepals, longer seeds with smaller, densely fibrillate appendage, and the habit of the inflorescence. Flowers of both species indicate autogamy but intermediates of unknown nature are known from Ecuador and Peru. 42. Luzula vulcanica Liebm., Vidensk. Meddel. Dansk Naturhist. Foren. Kjøbenhavn 2: 44 (1850) T: Mexico, Veracruz, Mt. Orizaba, 4250 m, F.Liebmann; lecto: C, fide H.Balslev, Fl. Neotrop. Monogr. 68: 65 (1996) [as ‘holo’]; isolecto: C, GH, HAL, K, P, S. Illustration: H.Balslev, Fl. Neotrop. Monogr. 68: 66, fig. 34, A, B (1996). Map: H.Balslev, Fl. Neotrop. Monogr. 68: 63, fig. 32 (1996). Perennials c. 10–45 cm tall, sparsely hairy; rhizome ascending to erect; stolons usually long, ascending. Stems erect, often slightly curved near the top (in the opposite direction to the nodding inflorescence). Basal leaves 3–15 cm long, usually 2–4 mm wide; cauline leaves usually 2, usually in upper half of stem, 1–5 (–11) cm long; all leaves acute, with papilloseserrulate margins. Bracts all scarious; lower bract membranous or castaneous (purplish), broadly lanceolate, c. 1 cm long, fimbriate-ciliate. Inflorescence a terminal spike like head, dark (reddish) castaneous brown, 1–4 × 0.5–1.2 cm, usually of 2–5 clusters, congested to slightly interrupted, suberect, if nodding then only the inflorescence not the upper part of stem. Bracteoles 1.5–2.0 mm long, usually castaneous-membranous. Tepals dark, reddishblackish, subequal, glabrous, usually 2.7–3.7 mm long, acuminate. Stamens 3; anthers 0.4–0.5 mm long; filaments 0.8–1.0 mm long; style 0.1–0.2 mm long; stigmas c. 0.5–0.8 mm long. Capsule oblong-ovoid, ±trigonous above, paler castaneous; capsule segments 1.6–1.9 × 79
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AUSTRALIAN BIOLOGICAL RESOURCES STU
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CONTENTS Contributors iv Species Pl
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Species Plantarum Steering Committe
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PREFACE This volume is the first of
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Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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