Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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SPECIES PLANTARUM — FLORA OF THE WORLD (2002)
Perennials (20–) 40–80 (–100) cm tall, loosely caespitose, subglabrous; rhizome creeping or
ascending, branched; stolons usually present, short or to c. 10 cm long. Leaves with densely
papillose-serrulate margins; apex acute; basal leaves few; cauline leaves 3–6 (upper 2 or 3
represent leaf-like bracts). Inflorescence usually lax, usually constituting 1/3– 1/2 of plant
height, decompound, storied; partial inflorescences subtended by leaf-like bracts; branches
diverging at wide angles; flowers borne singly, rarely in 2 (–3)-flowered clusters. Tepals
equal, glabrous, ±lanceolate, usually 1.8–2.8 mm long, pale reddish brown to dark
castaneous-brown. Stamens 6; anthers 0.5–1.1 mm long; filaments 0.6–1.0 mm long; style
usually 0.4–0.5 mm long; stigmas 1.0–1.5 mm long. Capsule obovoid to ±ellipsoid, subacute
to acute, with mucro 0.2–0.4 mm long, distinctly papillose above when ripe, equalling or
±longer than perianth; capsule segments c. 2.1–2.3 × 1.1–1.4 mm. Seeds deep brown,
1.2–1.4 mm long, (0.6–) 0.7–0.8 mm wide; apex usually distinct; basal appendage indistinct,
to 0.1 mm, fibrillate.
A widely distributed variable complex of populations extending from E Nepal to SW China,
Taiwan and S Malesia. The variation is centred in Sichuan and adjacent territories where two
varieties linked by occasional intermediates are recognized. The whole group is in need of
revision. Most common habitats include humid mountain forests, wet scrub, shady ravines
and cliff-faces.
Lower cauline leaves usually 4–11 mm wide; tepals ±brown to deep
(castaneous-)brown; stem usually less than 1.5–2.0 mm in diam. 23a. var. effusa
Lower cauline leaves usually 10–21 mm wide; tepals paler reddish brown to
brown; stem usually 3–6 mm in diam. 23b. var. chinensis
23a. Luzula effusa Buchenau var. effusa
Illustrations: Muh-Tsuen Kao & C.E.Devol, in Hui-Lin Li, Fl. Taiwan 5: 155, plate 1304 (1978); H.J.Noltie,
Fl. Bhutan 3(1): 263, fig. 25 (1994); K.F.Wu, Fl. Reipubl. Popularis Sin. 13(3): 240, plate 58 (1997).
Plants usually to (50–) 80 cm tall, often slender; stolons short to long. Stem usually 1–2 mm
diam. Cauline leaves usually 4–11 mm wide. Tepals ±brown to deep (castaneous-)brown,
usually 1.8–2.8 mm long. Anthers 0.5–1.1 mm long; filaments usually 0.6–0.8 (–1.0) mm long.
The East Himalaya from E Nepal to Manipur, N Burma, SW and C China, Taiwan, Luzon,
Borneo, Sulawesi and New Guinea. 36: CHC, CHN, CHT. 38: TAI. 40: EHM, NEP.
41: BMA. 42: BOR, NWG, PHI, SUL. Humid mountain woodlands, shrubby slopes and
subalpine ravines. Map 45.
36. TIBET: Kongbo, Doshong La, 4000 m, F.Ludlow, G.Sherriff & G.Taylor 4676a (BM, UPS). CHINA
SOUTH-CENTRAL: Sichuan, Mt Omei [Emei], Chiu-lo Tung, 2000 m, F.T.Wang 23312 (S). CHINA
NORTH-CENTRAL: Gansu ['Kansu orient.'], 4 Sep 1890, Potanin (K, LE). 38. TAIWAN: Taichung,
W slope of Mt Loyehwei-shan, Shimizu & Chuang 20183 (S). 40. NEPAL: Tamur Valley, Topke Gola,
c. 3530 m, J.D.A.Stainton 976 (UPS). BHUTAN: Pele-La, Gould 345 (K). 42. BORNEO: Sabah,
Kinabalu, Masilau R., M.S.Clemens 51371 (K). NEW GUINEA: Papua, Goilala, Mt Albert Edward, 3600 m,
J.R.Croft et al. 61394 (BM, K). PHILIPPINES: Luzon, Mt Pulog, E.D.Merrill 6490 (K). SULAWESI:
Enrekang, Rante Mario, 3300 m, P.J.Eyma 879 (K).
A very variable taxon. Rarely, very young plants have the inflorescence only indistinctly
storied. Plants from Taiwan and Malaya have tepals usually 2.5–2.8 mm long, longer than
capsules. The Malayan plants have more numerous basal leaves and, on average, longer
stolons. Intermediates between the two varieties are not rare.
23b. Luzula effusa var. chinensis (N.E.Br.) K.F.Wu, J. E. China Norm. Univ., Nat. Sci.
Ed., 1992(3): 92 (1992)
Luzula chinensis N.E.Br., J. Linn. Soc., Bot. 36: 161 (1903). T: China, [Sichuan] Szechuan, A.Henry 8829;
syn: K; S (fragment); China, Szechuan, near Tibetan frontier, Tachienlu [Kangding], A.Pratt 360; syn: BM, K.
Plants usually 50–100 cm tall, robust; stolons short. Stem usually 2–6 mm in diam. Cauline
leaves usually to 20 cm long and 10–21 mm wide. Tepals usually 1.9–2.3 mm long, paler
reddish brown to brown. Anthers 0.5–0.6 mm long; filaments 0.6–0.8 mm long.