Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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38 SPECIES PLANTARUM — FLORA OF THE WORLD (2002) long; stigmas 1.0–1.2 mm long. Capsule subglobular, ±obtuse, mucronate, castaneous-brown; capsule segments 1.7–2.0 mm long, including mucro 0.1–0.2 mm long and 1.4–1.5 mm wide. Seeds subglobular, pale castaneous-brown, smooth, shining, 0.8–0.9 (–1.0) mm long, 0.7–0.8 mm wide; apex pale; basal appendage indistinct, to 0.1 mm long, very sparsely fibrillate. 2n=12, fide N.Galland, Trav. Inst. Sci. Univ. Mohammed V, Sér. Bot., 35 (22): 106–121 (1988). Endemic to the Great Atlas and Anti-Atlas in Morocco. 20: MOR. Amongst shady mountain rocks in open oak woodlands. Map 31. 20. MOROCCO: Great Atlas, Oukaimeden, 2500–2600 m, P.H.Davis 67595 (E); Reraya, Sidi-Chamarouch, 2350–2500 m, 1923, R.V.Litardiére (MPU); Great Atlas, Reraya Valley, Sidi-Chamarouch, 2400–2500 m, 21 Jul 1924, R.Maire (P); Great Atlas, Gourdouze, 1800 m, Apr 1928, J.Braun-Blanquet & I.Braun-Blanquet (MPU). A species in many respects intermediate between sections Anthelaea and Diprophyllatae. The seed characters clearly point to Luzula (Marlenia) elegans Lowe. Sect. 3. NODULOSAE Luzula sect. Nodulosae Chrtek & Křísa, Preslia 46: 211 (1974) Luzula subsect. Nodulosae (Chrtek & Křísa) Novikov, Byull. Moskovsk. Obshch. Isp. Prir., Otd. Biol. 95(6): 66 (1990). T: Luzula nodulosa E.Mey. Perennials with thick nodulose horizontal rhizomes. Leaf tips acuminate, subulate. Lower bract ±membranous to subherbaceous. Inflorescence umbelloid or anthelate, composed of capituliform dense large clusters, on ±suberect peduncles. Seeds broadly ovoid, large, 1.7–1.9 × 1.3–1.4 mm, with apical and basal patches of looser outer seed-coat; appendages absent; base distinctly fibrillate. A monotypic section confined to the Mediterranean region, disjunctly in NW Africa and from Greece to W Anatolia. 12. Luzula nodulosa [Bory & Chaub.] E.Mey., Linnaea 22: 410 (1849), nom. cons. propos. Juncus nodulosus Bory & Chaub., Exp. Sci. Morée Bot. 3: 105, t. 11, f. 1 (1832), nom. illeg., non Wahlenb.; Juncus graecus Chaub. & Bory, Nouv. Fl. Pélop. 23, Plate 12, fig. 1 (1838); Luzula graeca (Chaub. & Bory) Kunth, Enum. Pl. 3: 310 (1841); Juncoides nodulosa (E.Mey.) Kuntze, Revis. Gen. Pl. 2: 724 (1891); Luzula nodulosa var. graeca (Chaub. & Bory) R.Maire, Bull. Soc. Hist. Nat. Afrique N. 22: 319 (1931). T: [Greece, Pylos, Arcadia] L.A.Chaubard; holo: P. Luzula nodulosa var. mauretanica R.Maire & Trab., in R.C.J.E.Maire, Bull. Soc. Hist. Nat. Afrique N. 22: 319 (1931). T: Algeria, Monts Tlemcen, entre Terni & Sebdou, L.Trabut; holo: P. Illustration: F.G.P.Buchenau, in H.G.A.Engler, Pflanzenr. (iv.36) 25: 65, fig. 46a, 66, fig. 46b (1906). Perennials c. (15–) 30–55 (–70) cm tall, loosely caespitose or one-stemmed; rhizome thick, c. 3–5 mm in diam., creeping to ascending, often with fibrous cover, nodulose, with internodes usually 0.5–2.0 cm long; stolons short, ascending. Lowermost leaves cataphyllous, c. 1–3 cm long, brownish; basal leaves flat, 7–16 (–20) cm long, 3.5–5.0 (–7.5) mm wide; cauline leaves 3–4 (–5), (1.5–) 2.5–5.0 (–15) cm long; margins densely ciliate, later sometimes glabrescent, densely papillose-serrulate; apex subulate. Lower bract brownish-membranous to subherbaceous, c. 1.0–3.5 (–5.0) cm long, shorter than inflorescence. Inflorescence of (1–) 2–5 (–15) large hemispherical to cup-shapedclusters; clusters (2–) 3–5 (–11)-flowered, 0.7–1.0 cm long, c. 1.0–1.5 cm wide; peduncles suberect, c. 2–10 cm long, smooth; secondary branches absent or frequent, distally inserted, short. Bracteoles scarious, often brownish at base, ovate, acuminate, c. 3.0–3.5 mm long; midrib distinct. Tepals ±equal, ovate-lanceolate or broadly lanceolate, c. 5.0–6.5 mm long, acute to acuminate; central band deep castaneous-brown; margins distinct, broad, membranous. Stamens 6; anthers 1.7–3.5 mm long, 2–10 times as long as filaments; filaments 0.3–0.9 mm long; style 1.8–2.7 mm long; stigmas c. 2.5–4.0 mm long. Capsule ovoid to broadly ovoid, acuminate, shorter than perianth, dark castaneous to pale castaneous-brown; capsule segments 3.3–4.0 × 2.3–2.8 mm
JUNCACEAE (Luzula) long; mucro to 0.5 mm long. Seeds broadly ovoid, deep brown, 1.7–1.9 mm long, 1.3–1.4 mm wide, with apical and basal whitish patches of loose outer seed-coat, distinctly fibrillate. Disjunctly in two areas: the E Mediterranean (from Greece to W Anatolia) and NW Africa (Algeria and Morocco). 13: GRC, KRI. 20: ALG, MOR. 34: EAI, TUR. Rocky slopes, open woodlands. Map 32. 13. GREECE: Attica, 650–1300 m, T.H.Heldreich [Herb. Graecum Norm.] 148 (P); Attica, Mt Pentelica, T.G.Orphanides 1105 (WU). KRITI: Askoyplicus, 1846, T.H.Heldreich (W); Hag. Vasilis, Kedros, K.H.Rechinger 849 (WU); Karpathos, Mt Kolla, W.Greuter 5314 (W). 20. ALGERIA: Kabylie, Mt Magris, 1700 m, E.Reverchon [Pl. Alger.] 364 (P, WU); Djurdjura, Ras Timedouine, 6 km E of Tikdja, 1860–1900 m, P.H.Davis 59408 (E). MOROCCO: Bab-Bou-Idir, C.G. van Steenis 19181 (L). 34. EAST AEGEAN IS.: Samos, Mt Ambelos, 800 m, K.H.Rechinger & F.Rechinger 3905 (K). TURKEY: Çanakkale, Erenköy, Renkoei Scala, 16 Apr 1883, P.E.E.Sintenis [It. Trojan.] 297 (W, UPS). Variable in stamen and pistil characters. There are two main forms, one with anthers 1.7–2.5 mm long, about 2–3 times as long as filaments, and style c. 2 mm long, the other with anthers 2.0–3.5 mm long, usually 4–8 times as long as filaments, and style c. 2.2–2.7 mm long. In N Africa, and in Morocco in particular, plants with longer anthers and styles predominate, mostly on limestone. Many African plants, however, are indistinguishable from the E Mediterranean ones, and intermediates are not rare. Sect. 4. DIPROPHYLLATAE Luzula sect. Diprophyllatae Satake, in T.Nakai & M.Honda, Nov. Fl. Jap. 1: 25 (1938) T: Luzula wahlenbergii Rupr.; lecto, designated here. Luzula ser. Effusae Satake, in T.Nakai & M.Honda, Nov. Fl. Jap. 1: 25 (1938); Luzula sect. Effusae [Satake] K.F.Wu, J. E. China Norm. Univ., Nat. Sci. Ed., 1992(3): 91 (1992). T: Luzula effusa Buchenau. Luzula ser. Spadiceae Satake in T.Nakai & M.Honda, Nov. Fl. Jap. 1: 26 (1938). T: Luzula wahlenbergii Rupr. Luzula sect. Glabratae Chrtek & Křísa, Preslia 46: 211 (1974). T: Luzula glabrata (Hoppe) Desv. Luzula subsect. Parviflorae Kovt., Bot. Žurn. (Moscow & Leningrad) 72: 1398 (1987); Luzula sect. Parviflorae [Kovt.] K.F.Wu, J. E. China Norm. Univ., Nat. Sci. Ed., 1992(3): 92 (1992). T: Luzula parviflora (Ehrh.) Desv. Perennials, subglabrous to sparsely ciliate; rhizome branched, ascending. Leaves with acute tips. Lower bract herbaceous to subherbaceous. Inflorescence decompound, a ±nodding to suberect panicle. Flowers usually borne singly, some in 2–5-flowered clusters. Tepals usually 2.0–2.5 (–3.5) mm long. Seeds usually ellipsoid to oblong-ellipsoid; appendage absent or, when visible, basal, fibrillate. A section with 12 species of mostly subarctic and oro-boreal, or alpine distribution. Main centres of diversity are in W North America and Europe; two species reach the southern hemisphere. Almost exclusively, diploids (2n=12 AL) and agmatoploids with 2n=24 BL are known in the section. The species are very close to one another in their general habit. 1 Upper part of capsule and mucro densely papillose; inflorescence usually storied (inflorescence subdivided in several subequal panicles separated by long rhachis segments; inflorescence usually constitutes 1/3 of plant height or more) 23. L. effusa 1: Capsule smooth; inflorescence storied or not so 2 Inflorescence storied 13. L. gigantea 2: Inflorescence not storied 3 Panicle branches abbreviated, and inflorescence therefore subcongested 16. L. subcapitata 3: Panicle branches (at least the main ones) long; panicle loose or subdense, never subcongested 39
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Page 3 and 4: CONTENTS Contributors iv Species Pl
Page 5 and 6: Species Plantarum Steering Committe
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