Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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SPECIES PLANTARUM — FLORA OF THE WORLD (2002)
Luzula spadicea var. candollei E.Mey., Linnaea 22: 401 (1849); Luzula spadicea subsp. candollei (E.Mey.)
K.Richt., Pl. Eur. 1: 185 (1890); Luzula alpinopilosa subsp. candollei (E.Mey.) Rothm., Feddes Repert. Spec.
Nov. Regni Veg. 67: 4 (1963). T: France, E Pyrenees, Val d’Eynes, 13 Sep 1839, R.J.Shuttleworth; syn: W;
Switzerland, Geneve, Castan; syn: n.v.; Pyrenees, Meles, L.Marchand; syn: n.v.
Illustrations: H.E.Hess, E.Landolt & R.Hirzel, Fl. Schweiz 1: 519 (1967), as L. spadicea; S.Pignatti, Fl. Ital. 3:
445 (1982).
Plants (10–) 15–20 (–35) cm tall. Leaves of sterile shoots (when present) usually 5–10 (–20)
cm long, 2–4 (–5) mm wide; cauline leaves usually 4–7 cm long, 1.5–3.5 (–5.5) mm wide.
Lower bracts subherbaceous or pale brownish, 2–3 (–4) cm long, substantially shorter than
inflorescence. Inflorescence usually 4–7 × 3–6 cm, suberect to slightly nodding, ±evenly
branched, most flowers usually borne singly, but with 2–4-flowered clusters usually also
present. Tepals 1.8–2.7 mm long. Anthers 0.9–1.2 mm long, usually 3–4 times longer than
filaments; filaments 0.3–0.4 mm long; style (0.7–) 0.9–1.2 mm long; stigmas c. 1.5 mm long.
2n=12 [AL], fide H.Nordenskiöld, Hereditas 37: 330–331 (1951); Druskovic, Int. Organ. Pl.
Biosyst. Newslett. 24: 13 (1995).
Mountains of Central and southern Europe: the Alps, the N Apennines, the Pyrenees, the
Vosges. 11: AUT, GER, SWI. 12: FRA, SPA. 13: ITA, YUG-SL. Rocky and stony subalpine
and alpine slopes. Map 47.
11. AUSTRIA: Steiermark, Bösenstein, 1896 m, G.Strobl [Fl. Exs. Austro-Hung.] 1872 (PR, PRC, WU);
Vorarlberg, Montafon, Hochmaderjoch, Gölles 462 (WU). SWITZERLAND: Tessin, St. Gotthard, Monte
Prosa, 2400 m, A.Kneucker [Cyper. Junc. Exs.] 90 (PR, PRA); Valais, Zermatt, Riffelberg, F.O.Wolf
[A.Kneucker, Cyper. Junc. Exs.] 90a (PR). 13. ITALY: N Apennines, Corno alle Scale, 21 Jul 1840, P.Savi
(PR); Saint-Rhémy, 2400 m, A.Charpin, W.Greuter & Hainard [Exs. Genav.] 117 (W).
Very variable but the two extremes (the broad-leaved, small-flowered form often recognized
as subsp. candollei, and the typical narrow-leaved large flowered plants) often coexist within
populations, and intermediates are not rare.
24b. Luzula alpinopilosa subsp. obscura S.E.Fröhner, Preslia 40: 426 (1968)
Luzula obscura (S.E.Fröhner) Novikov, Byull. Moskovsk. Obshch. Isp. Prir., Otd. Biol. 95(6): 66 (1990).
T: Slovakia, Vysoké Tatry, Zbojnická chata, 1900 m, 14 Aug 1965, S.Fröhner; holo: PR.
Luzula carpatica Kitt. ex Kanitz, Linnaea 32: 327 (1863), an nom. validum?; Luzula spadicea A.b. [unranked]
carpatica (Kitt. ex Kanitz) Asch. & Graebn., Syn. Mitteleur. Fl. 2(2): 513 (1904);. Luzula spadicea f.
carpatica (Kitt. ex Kanitz) I.Grint., Fl. Rep. Soc. Roman. 11: 594 (1966). T: [Slovakia], 'in der kleinen
Kohlbach', J.D.T.Mauksch; holo: n.v., probably BP.
Illustration: T.Szynal & J.Mądalski, Atlas Fl. Polsk. 2(2): 151 (1931).
Plants 10–30 cm tall. Cauline leaves 3–8 cm long, 3.5–6.5 mm wide. Lower bract usually
herbaceous, usually 1.5–4.5 cm long, shorter than inflorescence. Inflorescence c. 4–5 × 3–5 cm,
±nodding to suberect; main branches long; secondary branches often abbreviated; flowers
borne singly and usually in (2–) 3–5 (–7)-flowered loose clusters. Tepals (2.0–) 2.2–3.0 mm long.
Anthers 0.9–1.2 mm long, usually 4–9 times longer than filaments; filaments 0.1–0.2 (–0.3) mm
long; style 0.8–0.9 mm long; stigmas 1.2–2.0 mm long. 2n=12 [AL], fide J.Májovský &
A.Murín, Karyotax. Prehl. Fl. Slov. 361 (1987). Fig. 12.
C Europe, confined to the Carpathians from Slovakia and Poland to Romania. 11: CZE-SL,
POL. 13: ROM. 14: UKR. Map 48.
11. SLOVAKIA: Nízké Tatry, Králova Hola, 1700 m, 20 Aug 1898, K.Tocl (PR); Vysoké Tatry, Mlynická
dolina, 1700 m, 10 Jul 1925, K.H.Rechinger (W). 13. ROMANIA: Retezat [Mtns], Gura Apei, Zanoga Lake,
2 Jul 1970, J.Soják (PR); Muntii Fogarasului, Vistea, Sambatei Valley, 1200–2000 m, 30 Jun 1970, J.Soják
(PR). 14. UKRAINE: the East Carpathians, Mt Goverla, 2030 m, 7 Jul 1948, E.Bradis & A.Zanjatova
(MW); Zakarpatskaya area, Rachiv, Mt Turkul (1850 m) 16 km NE of Bogdan, 1740 m, 27 Jul 1996, Z.Kaplan
96/506 (PRA).
The overall variation of subsp. obscura is greater than that described in the protologue.
In particular, narrow-leaved and small-flowered plants occur in populations of subsp.
obscura. Some plants from Bulgaria (the Stara Planina Mtns) are intermediate between
subsp. obscura and subsp. deflexa.