Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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88 SPECIES PLANTARUM — FLORA OF THE WORLD (2002) mucronate, shorter than or equalling tepals; capsule segments ovate, 1.7–1.9 × 1.1–1.2 mm, dark brown above. Seeds brown to pale brown, ellipsoidal, 1.0–1.1 × 0.6 mm, apically darker, apiculate, basally fibrillate. 2n=36, 42, 48, cf. Dawe & Murray, Canad. J. Bot. 59: 1373–1374 (1981); H.Nordenskiöld, Bot. Not. 1949: 85–86 (1949). Fig. 20. Subarctic and mountain regions of northern Europe, with a NW limit in Iceland and reaching northern European Russia and Novaya Zemlya in the NE. Reported to occur in Greenland but material not seen. 10: FIN, FOR, GRB, ICE, NOR, SVA, SWE. 14: RUN. 30: WSB. Mountain and subarctic tundra and stony slopes. Map 77. 10: FØROYAR: Kunö Is., 24 Aug 1897, Hartz & C.E.Ostenfeld (LD, S). GREAT BRITAIN: Scotland, Easterness, Cairngorm, A.Melderis 425 (S). NORWAY: Dovre Fjeld, Fogstuen, 1000 m, 22 Jul 1889, K.G.Baenitz (W). SVALBARD: Safehaven, 30 Aug 1882, A.G.Nathorst s.n. (S); Jan Mayen, Wildberget, Lid [Fl. Exs. Ins. J.Mayen] 18 (LD). SWEDEN: Åsele Lappmark, Laxfjället, 12 Jul 1889, G.Halldin s.n. (LD); Lycksele Lappmark, Tärna socken, Brandsfjället, 24 Jul 1937, E.Asplund s.n. (LD); ‘ex alpibus Lapponiae Pitensis’ G.Wahlenberg s.n. (WU). 14. RUSSIA NORTH: Lapponia ponojensis, Ponoy, 27 Aug 1899, J.Montell s.n. (H). Sometimes difficult to distinguish from L. confusa in Scandinavia because of frequent occurrence of intermediates of hybrid origin 47b. Luzula arcuata subsp. unalaschkensis (Buchenau) Hultén, Ark. Bot. 7(1): 32 (1968) Luzula arcuata var. unalaschkensis Buchenau, Bot. Jahrb. Syst. 12: 124 (1890); Luzula unalaschkensis (Buchenau) Satake, in T.Nakai & M.Honda, Nov. Fl. Jap. 1: 31 (1938) & J. Jap. Bot. 14: 7 (1938). T: Alaska, Unalaschka, Mertens; lecto: W, designated here. [Unalaschka, J.F.G.Eschscholtz; orig. syn: LE, n.v.]. Luzula arcuata var. kamtschadalorum Sam., in E.Hultén, Kongl. Svenska Vetenskapsakad. Handl., ser. III, 5(1): 223 (1927); Luzula kamtschadalorum (Sam.) Gorodkov ex Krylov, Fl. Zap. Sibiri 3: 551 (1929); Luzula unalaschkensis subsp. kamtschdalorum (Sam.) Tolm., Arktic Fl. SSSR 4: 40 (1963). T: Russia, Kamchatka, Avatcha volcano, 20 Aug 1920, E.Hultén 670; lecto: S, fide L.Hämet-Ahti, Ann. Bot. Fenn. 10: 123 (1973); isolecto: K. Luzula beringensis Tolm., Arktic Fl. SSSR 4: 39 (1963). T: Russia, Terra Koriakorum, prope sinum Korfii maris Beringii, 19 Aug 1960, A.I.Tolmachev, Stepanova & Fedorova; holo: LE. Illustrations: E.Hultén, Fl. Alaska 299 (1968); L.Hämet-Ahti, Ann. Bot. Fenn. 10: 126, fig. 3; 127, fig. 4 (1973); V.S.Novikov, in S.S.Kharkevich, Sosud. Rast. Soviet. Dal. Vost. 1: 83, plate vi (1985). Maps: E.Hultén, Fl. Alaska 299 (1968); N.K.Kovtonyuk, in L.I.Malyshev & G.A.Peshkova, Fl. Sibiri 4 (Araceae – Orchidaceae): map 34 (1987) [Siberia]; J.Coffey Swab, Fl. North Amer. 22: 266 (2000). Perennials 10–25 cm tall; rhizome ascending; stolons ascending, usually 2–3 cm long. Leaves ±flat, sparsely ciliate, subabruptly narrowing to an acute tip; basal leaves usually 8–12 cm long, 3–4 (–5) mm wide; sheath base brown to purplish; cauline leaves 1–3, to 3–5 (rarely 8) cm long, 1.5–3.0 mm wide; sheaths sparsely ciliate above. Stem 0.3–0.4 (–0.5) mm diam. below inflorescence. Lower bract usually 5–8 (–15) mm long, brownish-membranous. Inflorescence usually corymbose with (2–) 6–10 (–17) glomerules; glomerules usually of (1–) 3–6 flowers each; peduncles capillary, arcuate, often nodding, often branched, usually reddish. Bracteoles ±ovate-lanceolate, c. 1.5 mm long, brownish to membranous, ciliate. Tepals ±equal, usually 2.0–3.0 × 0.6–0.8 mm, ovate to lanceolate, acuminate, often ciliolate, medium to pale brown. Stamens 6; anthers 0.4–0.6 mm long; filaments 0.4–0.5 mm long; style 0.3–0.4 (–0.5) mm long; stigmas 1.1–1.5 mm long. Capsule ellipsoidal with a short beak, shorter than tepals; capsule segments pale brown to brown, 1.9–2.2 × 1.0–1.4 mm. Seeds brown, narrowly ellipsoidal, 1.0–1.1 × c. 0.6 mm; appendage very short, up to 0.15 mm long; base fibrillate (on the top of caruncle). 2n=36, G.S.Knaben, Nytt Mag. Bot. 15: 244 (1968). Russian Far East, Japan, Baikal Lake region, northwestern America from Washington and British Columbia to Alaska and Nunavut [NW Territories]. Not confirmed but probably present in China and Mongolia. 30: BRY, CTA, IRK. 31: KAM, KUR, MAG. 38: JAP, KOR. 70: ALU, ASK, NWT, YUK. 71: ABT, BRC. 73: WAS. Rocky or gravelly snow-patches, heaths and slopes, often on bare ground. Map 78. 30: BURYATIYA: Baikal Range, sources of Kurkula R., T.V.Egorova & V.N.Siplivinsky s.n. (OULU). 31: MAGADAN: upper Main R., 12 Aug 1932, B.N.Gorodkov & B.A.Tikhomirov s.n. (S). KAMCHATKA: Natchika, Mt. Tschopotschka [Shapochka], 4 Aug 1908, V.L.Komarov s.n. (S). KURIL IS.: Shumushu,
JUNCACEAE (Luzula) G.Koidzumi 39053 (TNS). 38: JAPAN: Nagano, Daitenjo-Lake, J.Sugimoto 13437 (KYO). KOREA: Mt. Pai-tou-shan, T.Nakai s.n. (TI). 70: ALEUTIAN IS.: Unalaska, E.Hultén 6671 (S). ALASKA: White Pass, 20 Aug 1929, S.Enander s.n. (UPS). YUKON: Canol Road, pass between Teslin R. and Nisutlin R., A.E.Porsild & A.Breitung 10983 (S). 71: BRITISH COLUMBIA: Red Rose Tungsten Mine, 15 km E of Skeena Crossing, c. 1600 m, J.A.Calder et al. 15173 (S). A variable subspecies; in Asia, towards the southeast, a gradual increase in perianth and capsule size can be observed, and plants from Japan often have solitary flowers on peduncles. The Japanese plants have often been referred to as L. wahlenbergii; literature records under the latter name from NE China may also refer to subsp. unalaschkensis. 48. Luzula confusa Lindeb., Bot. Not. 1855: 9 (1855) Luzula arcuata f. confusa (Lindeb.) Kjellm., Vega-Exped. Vetenskapl. Iagttagelser 2: 59 (1883), n.v.; Luzula arcuata subsp. confusa (Lindeb.) O.C.Dahl, in A.G.Blytt, Handb. Norges Fl. 201 (1906); Juncoides confusa (Lindeb.) A.Heller, Cat. N. Amer. Pl., 3rd edn, 70 (1910). T: [Norway] Dovre, 1854, C.J.Lindeberg; lecto: S, designated here. Luzula spicata var. kjellmanii Nath., Kongl. Vetensk. Akad. Handl. 30: 28 (1884). T: N Greenland, Kap York, Insugigsok, 26 Jul 1883, A.G.Nathorst; syn: W. Luzula hyperborea R. Br., Chlor. Melvill. [Parry’s 1 st Voyage App.] 283 (1823), nom. rejic. prop.; Luzula arcuata var. hyperborea (R.Br.) Rink, in J.Reinhardt et al., Naturhist. Bidr. Grønl. 119 (1857); Juncoides hyperborea (R.Br.) Coville, Mem. Torrey Bot. Club 5: 107 (1894). T: Spitzbergen, W.Scoresby; syn: BM; Greenland, W.Scoresby; syn: BM; Melville Island, W.E.Parry; syn: BM, PH. [covering both Luzula confusa and Luzula nivalis]. Luzula confusa var. subspicata Lange, Consp. Fl. Groenland. 127 (1880); Luzula confusa f. subspicata (Lange) Krylov, Fl. Zap. Sibiri 3: 552 (1929). T: [Greenland] numerous syn: C. Luzula hyperborea var. extensa Scheutz, Kongl. Svenska Vetenskapsakad. Handl. 22: 169 (1888). T: [Russia, Taymyr] Nikandroviskij ostrov et Briochovskij ostrov; syn: n.v. Luzula confusa f. normalis Krylov, Fl. Zap. Sibiri 3: 552 (1929). T: [not indicated]. Illustrations: J.Lid, Norsk & Svensk Fl. 203, fig. 100b (1963); E.Hultén, Fl. Alaska 301 (1968). Maps: E.Hultén & M.Fries, Atlas N. Eur. Vasc. Pl. 1: map 169 (1986); E.Hultén, Fl. Alaska 301 (1968); N.K.Kovtonyuk, in L.I.Malyshev & G.A.Peshkova, Fl. Sibiri 4 (Araceae – Orchidaceae): map 35 (1987) [Siberia]. Perennials, usually 10–15 (–20) cm tall, sparsely hairy, loosely to densely caespitose with ascending densely branched to simple rhizome, sometimes with short ascending vegetative shoots; plant base usually deep purple-brown, shiny. Basal leaves up to 6–9 cm long, ±subcanaliculate to ±flat, 1.5–2.5 mm wide; tip acuminate; cauline leaves 1–2, usually 2–4 cm long, 1.0–1.5 mm wide. Stem 0.5–0.9 mm in diam. below inflorescence. Lower bract membranous, less often brownish-membranous, usually 4–9 mm long. Inflorescence composed of one sessile and often 1-2 pedunculate clusters; peduncles without secondary branches, erect, straight, less often arcuate or recurved, c. 0.3 mm in diam.; sessile cluster bigger (sometimes with a discernible basal subcluster), usually more than 10-flowered, 7–10 × 6–7 (–9) mm; pedunculate clusters smaller, 5–12 (–14)-flowered, 5–6 (–7) × 4–5 (–6) mm. Bracteoles ±ovate, brownish, c. 1 mm long; membranous apex fimbriate-ciliate. Tepals ±equal, usually 2.0–2.3 mm long, ±acuminate to acute, dark brown. Stamens 6; anthers 0.4–0.5 mm long; filaments 0.5–0.7 mm long; style c. 0.2 mm long; stigmas 0.8–1.0 mm long. Capsule ±ovate, with top rounded, ±not mucronate, shorter than or equalling tepals, blackish brown to dark castaneous brown; capsule segments ovate, 1.8–2.0 × c. 1.3–1.4 mm. Seeds narrowly ellipsoidal, 1.0–1.1 × 0.6 mm, ±pale brown, distally dark apiculate; base fibrillate; appendage not visible. 2n=36, Á.Löve & D.Löve, Univ. Colorado Stud. Ser. Biol. 24: 26 (1966). Fig. 19. Northernmost regions of Eurasia and North America, mountains of northern Asia. 10: FIN, ICE, NOR, SVA, SWE. 14: RUN. 30: ALT, BRY, CTA, IRK, KRA, TVA, WSB, YAK. 31: MAG. 37: MON. 70: ASK, GNL, NWT, YUK. 71: ABT, BRC. 72: QUE, NFL. 75: MAI, NWH, VER. Wet, exposed rocky and sandy hillsides, and mountain summits, tundra. Map 79. 10: NORWAY: Dovre, Knudshoe [Knutshøa], Aug 1875, H.G.Falk s.n. (LD). SVALBARD: Lina Red Bay, 14 Jul 1899, T.Wulff s.n. (W). SWEDEN: Torne Lappmark, Kiruna, 700 m, G.Simmons 121 (LD). 14. RUSSIA NORTH: Lapponia ponojensis, Orlov, A.O.Kihlman 907 (H). 30: WEST SIBERIA: Novaya Zemlya, 89
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AUSTRALIAN BIOLOGICAL RESOURCES STU
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CONTENTS Contributors iv Species Pl
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Species Plantarum Steering Committe
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PREFACE This volume is the first of
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