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Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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<strong>JUNCACEAE</strong> (<strong>Luzula</strong>)<br />

papillose-serrulate margins; apex acute. Lower bract herbaceous, usually 2.5–5 cm long,<br />

shorter than or almost equalling inflorescence. Inflorescence many-flowered, ±nodding, later<br />

suberect, usually 5–7 × 4–5 cm; flowers borne singly or crowded in 2 (3–4)-flowered groups.<br />

Bracteoles scarious, ovate, ciliate, c. 1.5 mm long. Tepals equal, glabrous, lanceolate,<br />

acuminate, usually 2.4–2.8 mm long, deep <strong>to</strong> dark castaneous-brown; margins inconspicuous,<br />

paler (straw-) brown. Stamens 6; anthers (1.0–) 1.1–1.4 (–1.7) mm long; filaments 0.3–0.5 mm<br />

long; style c. 0.8–1.0 mm long; stigmas 1.0–1.5 mm long. Capsule oblong-ovoid, ±acute,<br />

shortly mucronate, dark brown, shining, ±equalling, or slightly longer than perianth; capsule<br />

segments (2.4–) 2.5–2.8 (–3.0) × c. 1.3–1.5 mm. Seeds ±ovoid, 1.1–1.3 mm long, c. 0.8 mm<br />

wide, brown; appendage <strong>to</strong> 0.1 mm long, fibrillate. 2n=12 [AL], fide A.-M.Lambert &<br />

J.Giesi, Inform. Annuelles Caryosyst. Cy<strong>to</strong>gén. 1: 28 (1967).<br />

SW Europe, disjunctly in the E Pyrenees, Massif Central, Vosges and S Schwarzwald.<br />

Reported from Montes Cantábricos, N Spain, but material not seen [see P.Montserrat, Anales<br />

Inst. Bot. Cavanilles 21(2): 459-461 (1964)]. 11: GER. 12: FRA, SPA. Subalpine s<strong>to</strong>ny<br />

meadows, along streams, usually on non-calcareous substrates above 1200 m. Map 41.<br />

11. GERMANY: Schwarzwald, Belchen, Jul 1858, H.A. de Bary (W). 12. FRANCE: Vosges, Hohneck,<br />

I.S.Perrin [F.W.Schultz, Herb. Norm.] 1154 (PR); Puy-de-Dôme, Mont-Dore, Grande Cascade, 1200–1300 m,<br />

17 & 22 Jun 1971, Rastetter (H, LD); Auvergne, Puy-Mary, 1200 m, 28 Jul 1909, J.J.Héribaud (PR, W);<br />

Auvergne, Puy-Ferrand, 2 Sep 1852, E.Desvaux (H); Hautes Cévennes, S of Mt Mézenc, 1400 m,<br />

N.Y.Sandwith 3897 (K); E Pyrenees, Vallée d'Eyne, c. 2100 m, E.Wängsjö & G.Wängsjö 3366 (LD); E.M.Sennen<br />

[Pl. Esp.] 4055 (LD); E Pyrenees, L'Hospitalet, Ellman & N.Y.Sandwith 247 (K). SPAIN: E Pyrenees,<br />

Nuria, 1850 m, E.M.Sennen [Pl. Esp.] 1809 (BC, W).<br />

22. <strong>Luzula</strong> parviflora (Ehrh.) Desv., J. Bot. (Desvaux) 1: 144 (1808)<br />

Juncus parviflorus Ehrh., Beitr. Naturk. 6: 139 (1791); <strong>Luzula</strong> spadicea var. parviflora (Ehrh.) E.Mey.,<br />

Linnaea 22: 402 (1849); Juncoides spadicea var. parviflora (Ehrh.) Kuntze, Revis. Gen. Pl. 2: 724 (1891);<br />

Juncoides parviflora (Ehrh.) Coville, Contr. U.S. Natl. Herb. 4: 209 (1893). T: [Sweden, Lappland],<br />

C.Linnaeus; lec<strong>to</strong>: Flora Lapponica 124, Institut de France, Paris, designated here; pho<strong>to</strong> BM, PRA.<br />

<strong>Luzula</strong> parviflora var. densiflora Lange, Consp. Fl. Groenland. 125 (1880). T: [not indicated; numerous<br />

syntypes from E and W Greenland].<br />

<strong>Luzula</strong> parviflora var. sparsiflora Lange, Consp. Fl. Groenland. 125 (1880). T: [not indicated; numerous<br />

syntypes from Greenland].<br />

Map: E.Hultén & M.Fries, Atlas N. Eur. Vasc. Pl. 1: map 166 (1986).<br />

Perennials 30–90 cm tall, loosely caespi<strong>to</strong>se; rhizome ascending; s<strong>to</strong>lons short, ascending,<br />

less often absent. Basal leaves 10–20 cm long, usually 5–10 mm wide; cauline leaves (3–) 4–<br />

5, 3–10 mm wide. All leaves with papillose-serrulate margins; apex acute. Lower bract<br />

herbaceous or brownish membranous, usually 1.5–4.0 cm long, shorter than inflorescence.<br />

Inflorescence usually lax, sometimes nodding, with some main branches <strong>to</strong> 13 cm long, often<br />

arcuate; flowers borne singly. Bracteoles subglabrous <strong>to</strong> sparsely ciliate, ovate, membranous.<br />

Tepals equal, glabrous, ±lanceolate, 1.8–2.3 (–2.5) mm long. Stamens 6; anthers 0.3–0.6 mm<br />

long, as long as or shorter than filaments. Capsule smooth above. Seeds ±ellipsoid, brown <strong>to</strong><br />

dark brown, 1.0–1.4 mm long; base fibrillate.<br />

A widely distributed northern circumpolar species, most common in subarctic areas and in the<br />

mountains of N Europe, N Asia and temperate North America. Three subspecies are recognized.<br />

1 Inflorescence many-flowered with long main branches and short most<br />

distal secondary branches (peduncles); perianth segments medium brown 22a. subsp. parviflora<br />

1: Inflorescence few-flowered, with long main branches and long distal<br />

secondary branches (almost divaricate); perianth segments usually pale<br />

brown <strong>to</strong> stramineous<br />

2 Stem conspicuously reddish at base; cauline leaves mid-green, shining,<br />

tepals with a distinct midrib 22b. subsp. fastigiata<br />

2: Stem without reddish coloration or with indistinct reddish tinge at<br />

base; cauline leaves bluish or greyish green, dull, tepals with an<br />

indistinct midrib 22c. subsp. melanocarpa<br />

49

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