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Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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118<br />

SPECIES PLANTARUM — FLORA OF THE WORLD (2002)<br />

<strong>Luzula</strong> multiflora var. gracilis Buchenau ex V.I.Krecz., Žurn. Russk. Bot. Obshch. Akad.<br />

Nauk S.S.S.R. 12: 490 (1928).<br />

T: Russian Far East, Ayan, Avacha R. basin [not located at LE].<br />

Doubtful Hybrid<br />

<strong>Luzula</strong> ×winderiae Murr, Magyar Bot. Lapok 28: 67 (1930).<br />

T: [Austria] Vorarlberg, supra lacum Spullersee montis Arlberg, c. 1900 m, 7 Aug 1916, J.Murr; holo: n.v.<br />

The name was reported <strong>to</strong> belong <strong>to</strong> a hybrid between L. multiflora var. alpestris<br />

[=L. alpina] and L. spicata. The existence of the hybrid is very doubtful.<br />

PREDOMINANTLY AFRICAN SPECIES<br />

60. <strong>Luzula</strong> mannii (Buchenau) Kirschner & Cheek, Kew Bull. 55: 900 (2000)<br />

<strong>Luzula</strong> campestris var. mannii Buchenau, Bot. Jahrb. Syst. 12: 159 (1890); <strong>Luzula</strong> campestris subsp. mannii<br />

(Buchenau) Weim., Svensk Bot. Tidskr. 40: 173 (1946); isonyms: Sóo, Acta Bot. Acad. Sci. Hung. 16: 367 (1971);<br />

Fern.Casas, Fontqueria 33: 41 (1992). T: [Cameroon] Kamerun-Gebirge, 10000–13500 ft, Mann 2108;<br />

lec<strong>to</strong>: K; iso (fragment): W; Fernando Po, 8500', Mann 658, 1467; para K; Cameroons, H.H.Johns<strong>to</strong>n 48<br />

[the latter erroneously cited as Mann 48 by F.G.P.Buchenau]; para: K.<br />

Perennials 25–60 cm tall; rhizome horizontal or ascending; s<strong>to</strong>lons short, ascending. Leaves<br />

with obtuse tip and densely papillose-serrulate margins, hairy <strong>to</strong> densely hairy. Lower bract<br />

leaf-like, usually shorter than inflorescence. Inflorescence of mostly pedunculate clusters.<br />

Peduncles smooth, usually with short secondary branches. Capsule shorter than perianth.<br />

Seeds ovoid-subglobular, 0.6–0.8 mm wide; appendage almost invisible, less than 0.1 mm,<br />

hyaline.<br />

A rare endemic with three disjunct populations in tropical Africa (Equa<strong>to</strong>rial Guinea,<br />

Cameroon and the Uganda/Kenya border). Two subspecies are recognized.<br />

Anther/filament length ratio c. 1.5; peduncles ±straight and rigid, 0.3–0.5 mm<br />

in diam.; tepals blackish castaneous brown 60a. subsp. mannii<br />

Anther/filament length ratio 0.3–0.6; peduncles flexuose <strong>to</strong> arcuate, 0.2–0.3 mm<br />

in diam.; tepals pale castaneous brown <strong>to</strong> dark brown, usually with a narrow<br />

pale margin and often with a pale yellowish-stramineous middle part 60b. subsp. gracilis<br />

60a. <strong>Luzula</strong> mannii (Buchenau) Kirschner & Cheek subsp. mannii<br />

Plants 25–60 cm tall. Basal leaves usually 7–9 (–13) cm long, (4–) 5–6 (–8) mm wide;<br />

cauline leaves 2–3, usually 3.5–7.0 cm long and 2.5–4.5 mm wide. Stem rigid, erect, c. 1.7 mm in<br />

diam. Lower bract 2.0–<strong>6.</strong>5 cm long. Inflorescence composed of 3–4 ±sessile and<br />

2–7 pedunculate clusters; peduncles usually with secondary branches and 1–3 sessile clusters<br />

on each branch; peduncles straight, rigid, erect and patent (very rarely one of them<br />

±nodding), 0.3–0.5 mm in diam., 2–3 (rarely up <strong>to</strong> 11) cm long; clusters ±echinate,<br />

±globular, 0.6–0.9 (–1.2) cm in diam., usually with more than 13 flowers. Bracteoles long<br />

acuminate, membranous, often with cupreous middle part, up <strong>to</strong> 3 mm long, equalling<br />

perianth or shorter, densely ciliate. Tepals long acuminate, blackish castaneous brown,<br />

subequal, sometimes one or two outermost ones distinctly longer than inner ones, 2.6–3.0 mm<br />

long. Anthers usually 0.8–1.2 mm long; filaments 0.5–0.9 mm long; style 0.5 mm long;<br />

stigmas 1.5–1.8 mm long. Capsule ±globular, rounded above with an indistinct mucro;<br />

capsule segments 1.8–2.0 mm long and c. 1.5 mm wide, dark brown. Seeds 0.9–1.1 mm long,<br />

0.7–0.8 mm wide. 2n=42, fide J.K.Mor<strong>to</strong>n, Opera Bot. 121: 159–172 (1993), n.v. Fig. 2<strong>6.</strong>

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