Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

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44 SPECIES PLANTARUM — FLORA OF THE WORLD (2002) R.W.Holm & H.H.Iltis 485 (A, MO, P); Irazú, La Playita, 3300 m, Pettier 14125 (AAU). GUATEMALA: Huehuetenango, San Mateo Ixtatán, Sierra de los Cuchumatanes, 2900 m, D.E.Breedlove 11555 (DS, LL, MICH, US). In Panama and S Mexico, plants intermediate between L. denticulata and L. gigantea occur, but are rare. They are almost glabrous and have ±storied inflorescence (vouchers in AAU). Further study is necessary. 15. Luzula wahlenbergii Rupr., Fl. Samojed. Cisural. 58 (1845) T: North Russia, ‘Litt. austr. ins. Kolguev’, F.J.Ruprecht; syn: LE, UPS, W; Norway, Finnmark, Kaafjord, M.Blytt in E.Fries [Herb. Norm. fasc. VIII] 70; syn: UPS, S. Luzula spadicea var. kunthii E.Mey., Linnaea 22: 403 (1849). T: Sweden, Lapponia Pitensis, G.Wahlenberg; syn: W;. Norway, Finnmark, Kaafjord, M.Blytt in E.Fries [Herb. Norm. fasc. VIII] 70; syn: UPS, S. Luzula borealis Fr., Summa Veg. Scand. 219 (1846). T: ‘Per alpes Scandinaviae altioris totius’. Norway, Finnmark, Kaafjord, M.Blytt in E.Fries [Herb. Norm. VIII] 70; syn: UPS, S. Illustration: J. Lid, Norsk & Svensk Fl. 202, fig. 99a (1963). Maps: L.Hämet-Ahti, Aquilo, Ser. Bot., 3: 18, fig. 5 (1965); E.Hultén & M.Fries, Atlas N. Eur. Vasc. Pl. 1: map 167 (1986). Perennials c. 20–30 cm tall, subglabrous, caespitose, sometimes with short ascending vegetative shoots. Stem with several basal and 1–2 cauline flat leaves; basal leaves 5–10 cm × 3–5 mm; upper leaves up to 3–5 cm × 2–4 mm; leaf margins papillose-serrulate; leaf tip acute. Inflorescence a lax nodding ‘panicle’ of 10–30 flowers, c. 4 cm long and 3 cm wide; flowers borne singly, pedunculate, or in 2(–3)-flowered clusters. Lower bract 4–10 mm long. Bracteoles up to 2.0 mm long, fimbriate-ciliate near apex. Tepals equal, 2.0–2.8 mm long, ±acuminate, entire, ±brown. Stamens 6; anthers 0.4–0.6 mm long, ±equalling filaments; style up to 0.5 mm long; stigmas c. 1 mm long. Capsule ellipsoidal, ±subacuminate, ±exceeding tepals; capsule segments brown, 2.0–2.4 × 1.1–1.5 mm. Seeds narrowly ellipsoidal, 1.2–1.4 × c. 0.7 mm, dark castaneous brown, without appendage; base fibrillate. 2n=24 [BL], H.Nordenskiöld, Hereditas 37: 330–331 (1951). Arctic Eurasia and mountains of Scandinavia, Russian Far East and Transbaicalia, Alaska and Arctic Canada and Greenland. 10: FIN, NOR, SVA, SWE. 14: RUN. 30: BRY, CTA, KRA, WSB, YAK. 31: KAM, KHA, KUR, MAG, PRM. 70: ASK, GNL, NWT, YUK. 72: QUE. Arctic and alpine mossy tundra, wet grasslands. Map 35. 10. FINLAND: Lapponia enontekiensis, 69°N, Aug 1912, J.Montell s.n. (PR). NORWAY: Finnmark, Nord- Varanger, Fossefjellet, N.Lundquist 4944 (UPS). SWEDEN: Torne Lappmark, Jukkasjärvi, 24 Jul 1904, Ringselle s.n. (PR). 14. RUSSIA NORTH: Murmansk, Imandra Lake, R.R.Pohle 6980 (MW). 30. WEST SIBERIA: Tyumen, Distr. Priuralskiy, 2.5 km N of Shuchye. 31. RUSSIAN FAR EAST: Magadan Distr., Pitlekaj, 1879 F.Kjellman s.n. (S); Chukotka, Nedle, A.A.Krause 175 (UPS). Kamchatka, Pal’matkina R., 1932 B.N.Gorodkov & B.A.Tikhomirov s.n. (S). 70. ALASKA: Nunivak Is., Nash Harbour, 1938 J.P.Anderson 3878, 3875 (S); Brooks Range, De Long Mtns, Feniak Lake, 600 m, K.Holmen & O.Mårtensson 61-1130 (UPS). CANADA, NORTHWEST TERRITORIES [Nunavut]: Franklin Distr., Frobisher Bay, 1959, E.Hultén s.n. (S). Close to L. parviflora and L. piperi. From the former it can be distinguished by lower growth, narrower leaves, 1–2 cauline leaves; from the latter by linear-lanceolate cauline leaves and dark brown seeds (L.Hämet-Ahti, Aquilo, Ser. Bot., 3: 11–21, 1965). 16. Luzula subcapitata (Rydb.) H.D.Harr., Man. Pl. Colorado 151 (1954) Juncoides subcapitata Rydb., Bull. Torrey Bot. Club 1904: 401 (1904). T: USA, Colorado, Silver Plume, 1895, C.L.Shear 4611; holo: NY. Illustration: J.Coffey Swab, Fl. North Amer. 22: 262 (2000). Map: J.Coffey Swab, Fl. North Amer. 22: 261 (2000). Perennials 15–30 cm tall, loosely caespitose; rhizome ascending, with numerous short ascending vegetative shoots. Stem ±green. Basal leaves 1–4, 3.5–6.0 cm long, 4–5 mm wide; cauline leaves 3 (–4), with lower ones broader, 3–6 cm long, 2.0–5.5 mm wide. All leaves
JUNCACEAE (Luzula) apiculate, glabrous or with sparse hairs at sheath mouth. Lower bract up to 1.5 cm long, rarely 3 cm long, sometimes overtopping inflorescence. Inflorescence a single congested dark head composed of indistinct clusters; each flower usually with a short visible peduncle to 1 mm. Bracteoles 1.0 (–1.5) mm long, brown; apex obtuse, ciliate. Tepals equal, glabrous, ovate-lanceolate, 1.8–2.2 mm long, ±dark castaneous-brown; margins not visible. Stamens 6; anthers (0.3–) 0.4 mm long, slightly shorter than or equal to filaments; filaments c. 0.4 mm long; style 0.2–0.3 mm long; stigmas 0.8–1.0 mm long. Capsule subglobular-ovoid, equalling or slightly exceeding perianth, almost rounded above, with a short apiculate mucro; capsule segments 1.7–1.8 × 1.2–1.3 mm long. Seeds paler castaneous-brown, c. 1.0 mm long, c. 0.6 mm wide; appendage absent; base fibrillate. A restricted endemic of mountains in Colorado, USA. 73: COL. Damp, boggy areas near alpine lakes and streams, usually above 3000 m. Map 36. 73. USA, COLORADO: San Juan Co., Tenmile Basin, Animas R. drainage, SE of Silverton, Needle Mtn Quadrangle, J.Michener 789 (S); Willow Lake, W.W.Robbins 7933 (H). Luzula subcapitata occupies an isolated position in the section; it is the only species with abbreviated inflorescence branches and peduncles, i.e., with a ±congested inflorescence. 17. Luzula divaricata S.Watson, Proc. Amer. Acad. Arts (n.s.) 6: 302 (1879) Juncoides divaricata (S.Watson) Coville, Contr. U. S. Natl. Herb. 4: 209 (1893); Luzula spadicea var. divaricata (S.Watson) Buchenau, in H.G.A.Engler, Pflanzenr. (iv.36) 25: 63 (1906); Luzula parviflora subsp. divaricata (S.Watson) Hultén, Acta Univ. Lund. (n.s.) sect. 2, 39(2): 441 (1943); Luzula parviflora var. divaricata (S.Watson) Boivin, Naturaliste Canad. 94: 526 (1967). T: California, Sierras, 1863, W.H.Brewer 1794; syn: UC, fide L.Hämet-Ahti, Ann Bot. Fenn. 8: 369 (1971); W.H.Brewer 2069, 2334; syn: n.v.; Sierra Co., Summit Station, E.L.Greene; syn: n.v.; Sierra Co., near Castle Peak, J.G.Lemmon; syn: GH, n.v. Illustrations: W.L.Jepson, Fl. Calif. 1: 259, fig. 45c, d, e (1921); C.L.Hitchcock et al., Vasc. Pl. Pacif. Northw. 1: 216 (1969). Maps: L.Hämet-Ahti, Ann. Bot. Fenn. 8: 369, fig. 1 (1971); J.Coffey Swab, Fl. North Amer. 22: 258 (2000). Perennials (10–) 15–25 (30) cm tall, densely caespitose, pale green, subglabrous; rhizome short, ±ascending, subnodulose; stolons absent. Basal leaves to 20 cm long, 4–6 mm wide; cauline leaves 2–3, 3–5 mm wide. All leaves with papillose-serrulate margins; tip acute. Lower bract c. 1.0–2.5 cm long, purplish-stramineous to subherbaceous, much shorter than inflorescence. Inflorescence many-flowered, lax, c. 7–10 cm in diam., with rigidly divaricate branches (often at ±right angles); flowers borne singly; distal peduncles usually more than 1.0–1.5 cm long. Bracteoles ovate-lanceolate, membranous; base often pale brownish, c. 1.1–1.4 mm long, subglabrous. Tepals equal, glabrous, stramineous to pale brown, often with reddish tinge, ±lanceolate, long-acuminate, (1.8–) 2.0–2.2 (–2.4) mm long; apex often ±reflexed. Stamens 6; anthers 0.4–0.6 mm long; filaments c. 0.4–0.5 mm long; style 0.2–0.4 mm long; stigmas 0.7–1.5 mm long. Capsule ±oblong, usually pale brown, acute, exceeding perianth; capsule segments 1.9–2.2 × 1.0–1.2 mm long. Seeds brown, 1.0–1.2 mm long, c. 0.6 mm wide; appendage indistinct, to 0.1 mm, fibrillate. An endemic of the SW United States, only found in the Sierra Nevada in California and westernmost Nevada. 76: CAL, NEV. Reported to occur in Oregon but material supporting the literature records not seen. Subalpine forest to alpine granitic slopes; 2100–3700 m. Map 37. 76. USA, CALIFORNIA: Sierra Nevada, Mineral King, F.V.Coville & F.Funston 1542 (K); Butte Co., Lassen National Forest, Jonesville, Butte Ck, c. 1700 m, 19 Aug 1932, H.F.Copeland (H, PR); Fresno Co., Cirque Lake, C.H.Quibell & E.M.Quibell 7714 (W). NEVADA: Washoe Co., Mt Rose, 3100 m, A.A.Heller 10213 (K). 18. Luzula piperi (Coville) M.E.Jones, Bull. Montana State Univ., Biol. Ser. 15: 22 (1910) Juncoides piperi Coville, in C.V.Piper, Contr. U. S. Natl. Herb. 11: 185 (1906); Luzula wahlenbergii subsp. piperi (Coville) Hultén, Ark. Bot. 7(1): 33 (1968). T: USA, Washington, Okanogan Co., the Cascade Mtns, 1897, A.D.E.Elmer 678; holo: US. 45
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Page 3 and 4: CONTENTS Contributors iv Species Pl
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