Part 6. JUNCACEAE 1: Rostkovia to Luzula - Species Plantarum ...

JUNCACEAE (Luzula)
apiculate, glabrous or with sparse hairs at sheath mouth. Lower bract up to 1.5 cm long,
rarely 3 cm long, sometimes overtopping inflorescence. Inflorescence a single congested
dark head composed of indistinct clusters; each flower usually with a short visible peduncle to
1 mm. Bracteoles 1.0 (–1.5) mm long, brown; apex obtuse, ciliate. Tepals equal, glabrous,
ovate-lanceolate, 1.8–2.2 mm long, ±dark castaneous-brown; margins not visible. Stamens 6;
anthers (0.3–) 0.4 mm long, slightly shorter than or equal to filaments; filaments c. 0.4 mm
long; style 0.2–0.3 mm long; stigmas 0.8–1.0 mm long. Capsule subglobular-ovoid, equalling
or slightly exceeding perianth, almost rounded above, with a short apiculate mucro; capsule
segments 1.7–1.8 × 1.2–1.3 mm long. Seeds paler castaneous-brown, c. 1.0 mm long,
c. 0.6 mm wide; appendage absent; base fibrillate.
A restricted endemic of mountains in Colorado, USA. 73: COL. Damp, boggy areas near
alpine lakes and streams, usually above 3000 m. Map 36.
73. USA, COLORADO: San Juan Co., Tenmile Basin, Animas R. drainage, SE of Silverton, Needle Mtn
Quadrangle, J.Michener 789 (S); Willow Lake, W.W.Robbins 7933 (H).
Luzula subcapitata occupies an isolated position in the section; it is the only species with
abbreviated inflorescence branches and peduncles, i.e., with a ±congested inflorescence.
17. Luzula divaricata S.Watson, Proc. Amer. Acad. Arts (n.s.) 6: 302 (1879)
Juncoides divaricata (S.Watson) Coville, Contr. U. S. Natl. Herb. 4: 209 (1893); Luzula spadicea var.
divaricata (S.Watson) Buchenau, in H.G.A.Engler, Pflanzenr. (iv.36) 25: 63 (1906); Luzula parviflora subsp.
divaricata (S.Watson) Hultén, Acta Univ. Lund. (n.s.) sect. 2, 39(2): 441 (1943); Luzula parviflora var.
divaricata (S.Watson) Boivin, Naturaliste Canad. 94: 526 (1967). T: California, Sierras, 1863, W.H.Brewer
1794; syn: UC, fide L.Hämet-Ahti, Ann Bot. Fenn. 8: 369 (1971); W.H.Brewer 2069, 2334; syn: n.v.; Sierra
Co., Summit Station, E.L.Greene; syn: n.v.; Sierra Co., near Castle Peak, J.G.Lemmon; syn: GH, n.v.
Illustrations: W.L.Jepson, Fl. Calif. 1: 259, fig. 45c, d, e (1921); C.L.Hitchcock et al., Vasc. Pl. Pacif.
Northw. 1: 216 (1969).
Maps: L.Hämet-Ahti, Ann. Bot. Fenn. 8: 369, fig. 1 (1971); J.Coffey Swab, Fl. North Amer. 22: 258 (2000).
Perennials (10–) 15–25 (30) cm tall, densely caespitose, pale green, subglabrous; rhizome short,
±ascending, subnodulose; stolons absent. Basal leaves to 20 cm long, 4–6 mm wide; cauline
leaves 2–3, 3–5 mm wide. All leaves with papillose-serrulate margins; tip acute. Lower bract
c. 1.0–2.5 cm long, purplish-stramineous to subherbaceous, much shorter than inflorescence.
Inflorescence many-flowered, lax, c. 7–10 cm in diam., with rigidly divaricate branches
(often at ±right angles); flowers borne singly; distal peduncles usually more than 1.0–1.5 cm
long. Bracteoles ovate-lanceolate, membranous; base often pale brownish, c. 1.1–1.4 mm
long, subglabrous. Tepals equal, glabrous, stramineous to pale brown, often with reddish
tinge, ±lanceolate, long-acuminate, (1.8–) 2.0–2.2 (–2.4) mm long; apex often ±reflexed.
Stamens 6; anthers 0.4–0.6 mm long; filaments c. 0.4–0.5 mm long; style 0.2–0.4 mm long;
stigmas 0.7–1.5 mm long. Capsule ±oblong, usually pale brown, acute, exceeding perianth;
capsule segments 1.9–2.2 × 1.0–1.2 mm long. Seeds brown, 1.0–1.2 mm long, c. 0.6 mm
wide; appendage indistinct, to 0.1 mm, fibrillate.
An endemic of the SW United States, only found in the Sierra Nevada in California and
westernmost Nevada. 76: CAL, NEV. Reported to occur in Oregon but material supporting
the literature records not seen. Subalpine forest to alpine granitic slopes; 2100–3700 m.
Map 37.
76. USA, CALIFORNIA: Sierra Nevada, Mineral King, F.V.Coville & F.Funston 1542 (K); Butte Co., Lassen
National Forest, Jonesville, Butte Ck, c. 1700 m, 19 Aug 1932, H.F.Copeland (H, PR); Fresno Co., Cirque Lake,
C.H.Quibell & E.M.Quibell 7714 (W). NEVADA: Washoe Co., Mt Rose, 3100 m, A.A.Heller 10213 (K).
18. Luzula piperi (Coville) M.E.Jones, Bull. Montana State Univ., Biol. Ser. 15: 22
(1910)
Juncoides piperi Coville, in C.V.Piper, Contr. U. S. Natl. Herb. 11: 185 (1906); Luzula wahlenbergii subsp.
piperi (Coville) Hultén, Ark. Bot. 7(1): 33 (1968). T: USA, Washington, Okanogan Co., the Cascade Mtns,
1897, A.D.E.Elmer 678; holo: US.
45