103osa x pseudorusticana fR. arvensis x sty/asa).Plants in a hedge west <strong>of</strong> stile, south-west <strong>of</strong> Mudbrooks House (402.337), 1995, PW looklike R, arvensis but have very sparsely hairy leaves and conical discs, with variable fruits. Thisis <strong>the</strong> first record for East Sussex, confirmed A. L. Primavesi.*Rosa g/aucB. Red~leaved rose.Planted and spreading by seed and suckers on <strong>the</strong> edge <strong>of</strong> Maskett's Wood (428.285) to formlarge patches but scarcely naturalized.Only rarely recorded as naturalized in Britain (usually bird-sown from gardens). Native incentral Europe.Rosa sty/asa. Short-styled field-rose.This rose may be quite overlooked in Sussex and south-east England, and will no doubt befound elsewhere too in hedges and wood edges - check hairy, upright bushes for united stylesarising from a conical disc at <strong>the</strong> top <strong>of</strong> <strong>the</strong> fruit.We have seven sites mainly on <strong>the</strong> clays; one bush in hedge east <strong>of</strong> Monkshill Farm(394.339), 1995, AK; Toll Lane, one bush outside farm (459.263) and ano<strong>the</strong>r in hedge1452.266), 1995, TR; two bushes occur in a small unimproved field south-east <strong>of</strong> FairwarpFarm, 1993 +, PO 1473.265); several on south side <strong>of</strong> Furnace Wood (479.260), 1995, TR;single plant in woodland on roadside near Chuck Hatch (471.332), 1995, PO; one plant onwoodland edge by road near Fagot Stack Corner (488.324), 1995, PD. Also recorded fromtetrad 43G (Broadstone Warren - golf course area) in Briggs (1990) but not refound and an unexpected place for it.Mainly in sou<strong>the</strong>rn Britain north to <strong>the</strong> Midlands, and locally distributed in south and west Europe.Rosa canina. Dog-rose.The commonest rose on <strong>the</strong> <strong>Forest</strong> (as it is in Sussex and Britain) on wood edges, hedges andverges, but absent from heath areas. One plant in woodland near Priory Road was about 6metres tall.Glabrous roses with free styles are likely to be this species, though it is quite a variablespecies and <strong>of</strong>ten has a few small glands on <strong>the</strong> leaves and sometimes hairs on <strong>the</strong> midribsunderneath (Group Pubescentes = R. dumetorum auct. non Thuill.); <strong>the</strong>se hairy plants arequite widespread but have not been mapped separately.Widespread in Europe south <strong>of</strong> <strong>the</strong> Baltic, south-west Asia and North Africa.Rosa x dumetorum Thuill. (R. canina x obtusifolia).Three specimens <strong>of</strong> this hybrid collected by TR in 1995 (as R. obtusifolia or hairy R. canina !)have been named by A. L. Primavesi and are <strong>the</strong> first records for East Sussex: one bush bytelegraph pole in hedge on south side <strong>of</strong> track to Hunter's Farm (448.264); one on north side<strong>of</strong> old railway line, <strong>Forest</strong> Row (437.348); several in a hedge at Quabrook 1444.343). Thishybrid is quite frequent where <strong>the</strong> parents grow near one ano<strong>the</strong>r, and it shows a bewilderinggradation <strong>of</strong> forms between <strong>the</strong> parents.Rosa obtusifolia. Round-leaved dog-rose.Recorded from near Crow borough Warren by Wolley-Dod (1937) and possibly still around but not refound. It is said to befrequent in sou<strong>the</strong>rn Britain in hedgerows and scrub, but we have very few recent records in Sussex. It is also found incentral, sou<strong>the</strong>rn and north-west Europe.
104Rosa tomentosa. Harsh downy-rose.This species, our only member <strong>of</strong> <strong>the</strong> downy-rose group, has large, pointed, quite hairy [eaves I• • which look grey in colour, but no significant glands except on <strong>the</strong> peduncles. It is usually a• hedgerow species which can grow to 4 metres, but also grows in scrub and on woodlandedges, It occurs widely in sou<strong>the</strong>rn Britain and is locally common on <strong>the</strong> chalk.Rare but quite characteristic <strong>of</strong> hedges around <strong>the</strong> edges <strong>of</strong> <strong>the</strong> <strong>Flora</strong> area, especially on<strong>the</strong> north-east side <strong>of</strong> <strong>the</strong> <strong>Forest</strong>. One plant outside garden, Priory Road (421.3481; west <strong>of</strong>Prickets Hatch (442.2701; at least four sites around Quabrook and Parrock Lane (44.341; Iwest end <strong>of</strong> Cackle Street (45.26); a few bushes in lane north <strong>of</strong> Hart's Farm (459.3351;somewhere near Newnham (49.28); one plant in hedge by junction at Hoadley's Farm!(507.325). Not yet refound in tetrad 42U where it was recorded in Hall (1980).It occurs mainly in sou<strong>the</strong>rn Britain but reaching Scotland, widespread in Europe except <strong>the</strong> far north, Caucasus and IAsia Minor.Rosa rubiginosa. Sweet~briar.One bush on east verge <strong>of</strong> A26 near Pound gate (486.285), 1994, TR et al. It was mown flat iin autumn 1995, but will probably re~grow.The sweet~briar group (R. rubiginosa and R. micranthal are distinguished by <strong>the</strong> apple~scented, long~stalked glands on <strong>the</strong> leaves, and <strong>the</strong>y can sometimes be found by smell fromacross <strong>the</strong> road! R. rubiginosa is distinguished from R. micrantha by <strong>the</strong> hairy styles, and <strong>the</strong>y,also differ in habit, sepal and pedicel characters.This rose is most characteristic <strong>of</strong> chalk soils in sou<strong>the</strong>rn Britain (it grows on clays atPoundgate), but also occurs in hedges and on suitable soils elsewhere north to Scotland.Widespread in Europe south <strong>of</strong> <strong>the</strong> Baltic, and also found in western Asia.Rosa micrantha. Small~flowered sweet~briar.Nutley, N. J. Treutler (Wolley-Dod 1937).• Scattered around, usually as single bushes, but rarely recorded from hedges. Northverge west <strong>of</strong> Wych Cross Place (418.319), TR, 1995; one bush at entrance to Birches onRidge Road (445.329), 1995, PW, one by pond and one on road to cricket ground nearby;hedge, Quabrook (444.343), 1995, TR; one on wood edge at Newbridge (459.326), 1995,PW (ano<strong>the</strong>r in hedge to east, not very glandular and possibly hybrid but hips trimmed <strong>of</strong>f,1995, TR); bank opposite Fairwarp turning (465.264), 1995, TR; Ridge Road west <strong>of</strong><strong>Ashdown</strong> Park Hotel (426.324), 1995 TR & PW; one just inside entrance to Crows (or Doves)Nest (474.283), 1994, <strong>Flora</strong> meeting; one enormous bush c. 5 metres tall on track opposite<strong>Forest</strong> Lodge Farm (497.327), 1987-1995, PW; one near Fisher's Gate (493.336), 1987-1995, PW; one on roadside nearSt John's Church (502.320), 1995, TR;Plants tend to have small leaves with many glands, strongly curved prickles, and glabrous styles. It seems mostfrequent in <strong>the</strong> eastern half <strong>of</strong> <strong>the</strong> <strong>Forest</strong> for no reason obvious as yet. It is commonest on chalky soils in sou<strong>the</strong>rn Britain,but is scattered through <strong>the</strong> Weald. Widespread in south, central and western Europe.*Prunus persica. Peach.One small shrub in bramble bush at Churl wood car park (416.310), 1993, <strong>Flora</strong> meeting,presumably resulting from a stone discarded after a picnic.Occasionally recorded in Britain elsewhere as a throw~out. It probably originally camefrom China.*Prunus cerasifera. Cherry plum.,--,--===----,--, Smockfarthing area (39.32), 1995, AK; one shrub in hedge on Parrock Lane (446.343), 1995,TR & PA; one small tree in hedge opposite entrance to Boringwheel Mill Farm (456.264),• 1995, TR & PA; Friar's Gate 149.33), 1995, SR.The first species to flower in spring (excepting <strong>the</strong> flowering cherries in gardens), whichis <strong>the</strong> easiest time to find it, but a search <strong>of</strong> roadside hedges in March 1995 failed to revealany more. Non~flowering specimens in shade could also occur. No fruit was seen in 1995; itapparently fruits rarely and ra<strong>the</strong>r sparingly in Britain.Mainly planted in hedges but also spread by birds, and increasing in England (Rich &Woodruff 1996). Recorded occasionally in West Sussex, probably under-recorded in EastSussex, and widespread in Britain. Native in south-east Europe and south~west Asia.
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Flora of Ashdown Forest,-,;I
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Copyright Tim Rich, Pat Donovan, Pa
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ACKNOWLEDGEMENTSWe would like to th
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THE HABITATS OF ASH DOWN FOREST3His
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Near the bottom of all the non~wood
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Conopodium majusCrataegus laevigata
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Fincham (1995) published a small ph
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13ANAL VS IS OF THE FLORA DATAIn th
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15The number of different visits to
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17Hedgerows Disturbed/waste ground
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19The frequencies with which specie
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21(r~0.434, pO.l).-0.059,•300t350
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23A second type of comparison has b
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25Similarly, environmental variable
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SPECIES ACCOUNTS27Lat'1n names foll
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29EQUISETACEAEEquisetum f/uviatile.
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One plant on a vertical, north-faci
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Viola pa/ustris in an open alder co
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35DRYOPTERIDACEAEPolystichum aculea
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37BLECHNACEAEBlechnum spicant. Hard
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39AngiospermsLAURACEAE* Laurus nobi
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41Ranunculus repens, Creeping butte
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wet mud, small streams, ditches and
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45*Meconopsis cambrica. Welsh poppy
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47Ulmusx hollandica f?U. glabra x m
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49FAGACEAEFagus sylvatica. Beech.Fo
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A special large oak at Chuck Hatch,
- Page 58 and 59: Cory/us avel/ana. Hazel.Recorded in
- Page 60 and 61: plants in the autumn; we have not r
- Page 62 and 63: Sagina procumbens. Procumbent pearl
- Page 64 and 65: 59*Saponaria officinalis. Soapwort,
- Page 66 and 67: 61Dolygonum arenastrum. Equal-leave
- Page 68 and 69: 63Rumex conglomeratus. Clustered do
- Page 70 and 71: 65Hypericum humifusum. Trailing S1
- Page 72 and 73: 67DROSERACEAEDrosera rotundifolia.
- Page 74 and 75: 69Viola laetea. Pale dog-violet.Thi
- Page 76 and 77: 71* Salix alba. White willow.3 -+-t
- Page 78 and 79: 73Salix repens var. repens (var. er
- Page 80 and 81: 75*Rorippa sylvestris. Creeping yel
- Page 82 and 83: 77* Thlaspi arvense. Field pennycre
- Page 84 and 85: 79aphanus raphanistrum. Wild radish
- Page 86 and 87: ica tetralix. Cross~leaved heath.As
- Page 88 and 89: centre of the Forest on alluvial so
- Page 90 and 91: It is predominantly a plant which o
- Page 92 and 93: 87SAXIFRAGACEAE[*Darmera pe/tata. I
- Page 94 and 95: 89insipid in flavour IHaskel1 1960)
- Page 96 and 97: 91ection Rubus. Subsection Hiemales
- Page 98 and 99: 93Rubus pr%ngatus.Nutley (45.27), 1
- Page 100 and 101: 95bus vestitus.Newbridge, 1969, B.
- Page 102 and 103: 97Rubus largificus.3 -+t--------'+_
- Page 104 and 105: Potentilla anserina. Silverweed.Com
- Page 106 and 107: 101Geum urbanum. Wood avens.Primari
- Page 110 and 111: 105Prunus spinosa. Blackthorn, Wint
- Page 112 and 113: orbus aucuparia. Rowan, Mountain as
- Page 114 and 115: 109r*Crataegus persimilis. Broad-le
- Page 116 and 117: 111icia tetrasperma. Smooth tare.Fo
- Page 118 and 119: 113Trifolium rep ens. White clover"
- Page 120 and 121: 115Genista tinctoria. Dyer's greenw
- Page 122 and 123: *Genista anglica. Petty whin, Needl
- Page 124 and 125: Myriophyllum alterniflorum. Alterna
- Page 126 and 127: 121Epilobium montanum x obscurum.Fo
- Page 128 and 129: habitats.It lives up to the name of
- Page 130 and 131: 125,UPHORBIACEAEVlercurialis perenn
- Page 132 and 133: Although one of Britain's tiniest p
- Page 134 and 135: 129*Oxalis articulata. Pink-sorrel.
- Page 136 and 137: *Erodium cicutarium. Common stork's
- Page 138 and 139: Widespread in Europe from the shore
- Page 140 and 141: * Sison amomum. Stone parsley.Tetra
- Page 142 and 143: 137Centaurium pu/chellum. Lesser ce
- Page 144 and 145: 139CONVOLVULACEAEConvolvulus arvens
- Page 146 and 147: 141*Nymphoides pe/tata. Fringed wat
- Page 148 and 149: Myosotis secunda (M. repens). Creep
- Page 150 and 151: 145Stachys 8lvensis. Field woundwor
- Page 152 and 153: 5cutellaria x hybrida (5. galericul
- Page 154 and 155: Origanum vu/gare. Marjoram.Near Fis
- Page 156 and 157: CALLlTRICHACEAE151Ca/litriche is a
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153Fraxinus excelsior. Ash.4 5Local
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~'Cymbalaria mural is. Ivy-leaved t
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157Veronica scutellata. Marsh speed
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159Sibthorpia europaea. Cornish mon
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161OROBANCHACEAEOrobanche rapum~gen
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163RUBIACEAESherardia arvensis. Mad
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165Viburnum OpUIU5.Guelder-rose, St
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167Valeriana officina/is 5.1. Commo
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169Cirsium vu/gate. Spear thistle.W
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171We suspect that Leontodon hispid
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173Mycelis muralis. Wall lettuce.Se
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175Taraxacum exceJlens.Scattered al
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177Taraxacum sinuatum.Recorded from
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179Crepis capillaris. Smooth hawk'
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181Hieracium. Hawkweeds.3134 5Hiera
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*So/idago gigantea. Early goldenrod
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ecorded with Chamaemelum are Matric
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*Senecio x albescens (S. cineraria
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*Petasites japonicus. Giant butterb
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*E1odea canadensis. Canadian waterw
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Arum maculatum. Lords~and~ladies, C
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Juncus x surrejanus (J. acutiflorus
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fences or topography. It may domina
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Eleocharis multicaulis. Many-stalke
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A hybrid first described by Col em
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203Carex riparia. Greater pond~sedg
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Material which initially looked lik
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207Festuca pratensis. Meadow fescue
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209Cynosurus cristatus. Crested dog
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211Glyceria f/uitans. Floating swee
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213Deschampsia cespitosa. Tufted ha
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Agrostis gigantea. Black bent.Tetra
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217Bromus lepidus. Slender soft~bro
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* Triticum aestivum. Bread wheat.A
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221L1L1ACEAENarthecium ossifragum.
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223*Allium triquetrum. Three-Gornar
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Trumpet length(mm)454035302520•
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227ORCHIDACEAEEpipactis purpurata.
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een unable to refind any bog orchid
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231DactylorhizB praetermissa. South
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Rachel A. Nicholson (1934-). Rachel
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235REFERENCESAbbas, M. H. (1979). S
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Graham, G. G. & Primavesi, A. L. (1
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Rich, T. C. G. & Smith, P. A. (1996
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GAZETTEER241This gazetteer has been
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Smockfarthing 40.31Smuggler's car p
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Asple adi+cuttr int~~~tl·Bteel SI!
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Broad bean 111 Cardamine x hausskne
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Fen bedstraw 163 Fumitory 46 Greate
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Leucanthemum maximum 186 Many-flowe
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Quercus ilex 50 Rorippa x ster/lis
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Sticky mouse-ear 56 Thale cress 73