Eleocharis multicaulis. Many-stalked spike-rush_Bogs on <strong>Ashdown</strong> <strong>Forest</strong> (Coleman 18361. <strong>Ashdown</strong> <strong>Forest</strong>, 1895, T. Hilton (BTN).<strong>Forest</strong> Row and <strong>Ashdown</strong> <strong>Forest</strong>, W. Borrer (Wollev-Dod 19371. Near Nutlev, 1957, R.A. Boniface (BRC). Tetrads 42J, 42N, 43F and 43M (Hall 1980).Very local on <strong>the</strong> wet heathy and peaty parts <strong>of</strong> <strong>Ashdown</strong> Fores( usually on soilskept open by trampling or disturbance. It also grows on <strong>the</strong> edges <strong>of</strong> <strong>the</strong> smallheathland ponds in Pippingford Park. It is <strong>of</strong>ten associated with Drosera spp. and4 5Rhynchospora.Locally distributed in Britain, most commonly in western Scotland. A calcifuge <strong>of</strong>western North Africa and predominantly western Europe.Scirpus sylvaticus. Wood clubrush.Tetrads 42N, 42T, 43L, 43M and 43R (Hall 1980).Locallv frequent on <strong>the</strong> north side <strong>of</strong> <strong>the</strong> <strong>Forest</strong> (as found bV Hall) usuallv in dampflushes, ponds and by streams and rivers. It is quite palatable and <strong>the</strong> flowering headsare <strong>of</strong>ten eaten <strong>of</strong>f, presumably by deer.3 Scattered in lowland Britain with <strong>the</strong> main headquarters being <strong>the</strong> Weald, where it iswidespread. Most <strong>of</strong> Europe but rare in <strong>the</strong> Mediterranean region.1994 5*Schoenoplectus lacustris (Scirpus lacustris). Common club-rush.One clump planted in pond in a Pippingford Park (444.303), PW. 1995.Locally distributed in Sussex but nearly absent from <strong>the</strong> High Weald. An emergentaquatic <strong>of</strong> ponds, rivers and lakes throughout Britain and Europe, Widespread in <strong>the</strong>world,3 -1-+-----''"-------'--1-4 5Isolepis setacea (Scirpus setaceus). Bristle club-rush.Bogs on <strong>Ashdown</strong> <strong>Forest</strong> (Coleman 1836). On <strong>the</strong> <strong>Forest</strong>s, E. Jenner (Arnold 1887),<strong>Ashdown</strong> <strong>Forest</strong>, 1901, T. Hilton (BTN; Wollev-Dod 19371. Five tetrads on <strong>the</strong> <strong>Forest</strong>(Hall 1980).Very locally scattered on rides and in open,. damp flushes; <strong>the</strong>re may be morearound but it is so inconspicuous we would not be surprised if it has been overlooked,This species is widespread and increasing in Britain (Rich & Woodruff 1990), Most<strong>of</strong> Europe except <strong>the</strong> north-east, temperate Asia and North Africa.4 5Eleogiton f/uitans (Scirpus f/uitans). Floating spike-rush.Bogs on <strong>Ashdown</strong> <strong>Forest</strong> (Coleman 1836). Pond on <strong>the</strong> <strong>Forest</strong>, near Wvch Cross(Whitwell 1902). <strong>Ashdown</strong> <strong>Forest</strong> near Nutlev (Done 19141. Chelwood Common, MissM. Cobbe (Wollev-Dod 1937). Near Nutlev, 1957, R. A. Boniface (BRCI. Five tetrads on<strong>the</strong> <strong>Forest</strong> (Hall 1980).3 It was found in nine 1-km squares during our survey, It is sometimes abundant, asin a small pond near Goat crossroads (401.326), 1995, AK, where many hundreds <strong>of</strong>plants were present in very shallow water and on wet mud as <strong>the</strong> water level fell in <strong>the</strong>4 5 summer heat. S'lmilar behaviour was seen in a pond in Kidbrooke Park (418.339), 1995,TR. A patch in <strong>the</strong> training pond on Pippingford Park (455.310),1987, SBRS, did notlast for long when <strong>the</strong> Army started training in it!It is a plant <strong>of</strong> base-poor water and is ra<strong>the</strong>r uncommon nationally, occurring mainly in <strong>the</strong> north and west andaround <strong>the</strong> New <strong>Forest</strong> but with a number <strong>of</strong> sites in Sussex and Surrey. Because <strong>of</strong> its grass-like appearance it canbe easily overlooked but <strong>the</strong> ra<strong>the</strong>r characteristic pale green colour and somewhat limp appearance allow it to bepicked out once it is known.The distribution in Sussex is concentrated around <strong>Ashdown</strong> <strong>Forest</strong> and south <strong>of</strong> Crawley. Widespread in Britainbut rare in <strong>the</strong> south-east. Western and western-central Europe.
200*Cyperus longus. Galingale.A couple <strong>of</strong> plants introduced with o<strong>the</strong>r aquatics to a pond on <strong>the</strong> golf course, <strong>Forest</strong>Row (436.3421 and ra<strong>the</strong>r eaten by deer, 14 August 1995, TR. The very next daydiscovered quite independently by PW I just as a bulldozer was clearing out <strong>the</strong> pond; it3 -+-1-----'--1-survived, but looked even worse for wear.As a native this plant is confined to a few marshes and wet pastures near <strong>the</strong> southand west coasts between Anglesey and Chichester. It is widely available from garden4 5centres and has been introduced to many ponds in sou<strong>the</strong>rn England (Stewart, Pearman& Preston 1994), It is native in sou<strong>the</strong>rn, central and western Europe, north to sou<strong>the</strong>rnEngland. It also occurs in Asia and North Africa.Rhynchospora alba. White beak-sedge.<strong>Ashdown</strong> <strong>Forest</strong>, 1805, E. Forster (Wolley-Dod 19371. On all <strong>the</strong> bogs <strong>of</strong> <strong>Ashdown</strong><strong>Forest</strong> (Forster 18161. On <strong>the</strong> great bogs <strong>of</strong> <strong>Ashdown</strong> (Cooper 18351. <strong>Forest</strong> Bogs on<strong>Ashdown</strong> <strong>Forest</strong> (Coleman 18361. Near Mareslield, 1852, W. C. Unwin (Wolley-Dod19371. Near <strong>Forest</strong> Row (Deakin 18711. <strong>Ashdown</strong> <strong>Forest</strong>, W. Borrer (Arnold 18871.3 -++-''"-:..-I--r-l- <strong>Ashdown</strong> <strong>Forest</strong> near Nutley, July 1889, Nutley, Dr Trentler (BTNI. <strong>Ashdown</strong> <strong>Forest</strong>,4 51901, T. Hilton (BTNI. <strong>Ashdown</strong> <strong>Forest</strong> (Whitwell 1902). Bog on <strong>Ashdown</strong> <strong>Forest</strong> nearLutby (Done 19141. <strong>Ashdown</strong> <strong>Forest</strong>, 1932, L. M. Child (TLSI. Bog near DuddleswellToll Gate, E. Jenner; near <strong>Forest</strong> Row, E. Jenner; Chuck Hatch, rare, Miss K, Pickard;<strong>Ashdown</strong> <strong>Forest</strong>, A. H. Wolley-Dod (Wolley-Dod 19371. Frequent in bogs at Kidbrookeand east <strong>of</strong> Broadstone Warren, 1948, and near Nutley, 1957, R. A. Bonilace (BRCI. South <strong>of</strong> Chuck Hatch, 1950s,FR. Frequent on <strong>the</strong> <strong>Forest</strong> (Hall 1980), <strong>the</strong> only East Sussex sites.Very restricted and localised on <strong>the</strong> centre <strong>of</strong> <strong>the</strong> <strong>Forest</strong>, usually in wet, acid peaty places where <strong>the</strong> peat islightly disturbed. It <strong>of</strong>ten grows with Drosera spp. and Eleocharis multicaulis. The sites are as follows: hundreds <strong>of</strong>plants on trampled ride east <strong>of</strong> Isle <strong>of</strong> Thorns (423.303), 1993, AK; ride north-east <strong>of</strong> Millbrook Farm (438.292),1994, PW; Millbrook (442.2931, 1993, AFRR; flushes north-east <strong>of</strong> <strong>Ashdown</strong> <strong>Forest</strong> Centre (439.329, 441.327,441.325,443.3281, 1993-1995, NM, AK, PW, etc.; Misbourne, two patches in valley bog at 451.275, and at455.278 and 457.274,1993+, PW, NM et al.; west <strong>of</strong> Ellison's Pond (461.289), 1986, PW; small patch in bog andodd ones near pond east <strong>of</strong> Old Lodge (468.2981 and to north (465.301, 468.3041, 1988-1993 +, PW; north <strong>of</strong>Poundgate (478.288 to 480.2871, 1993, AFRR; boggy area by ride south <strong>of</strong> New Pond Cottages (481.293), 1995,RN & ER. Also seen at 422.307,443.332,439.291 and 449.289, 1990-1991, FR.Very rare in West Sussex on <strong>the</strong> Lower Greensand but common in <strong>the</strong> New <strong>Forest</strong> and at Wool mer in Hampshire.Locally distributed in Britain, most common in western Scotland, Most <strong>of</strong> Europe except <strong>the</strong> Mediterranean region and<strong>the</strong> south-east, and found in North America.Carex. Sedges.Full details <strong>of</strong> this large genus can be found in <strong>the</strong> BSBI Sedge handbook (Jermy et al. 19821.Carex paniculata. Greater tussock-sedge.In some <strong>of</strong> <strong>the</strong> bogs on <strong>Ashdown</strong> <strong>Forest</strong> (Coleman 1836), Plentiful in a bog south <strong>of</strong>Coleman's Hatch (Done 19141. <strong>Ashdown</strong> <strong>Forest</strong>, Miss K. Pickard (Wolley-Dod 1937).Tetrads 42N, 43L, 43M and 43R (Hall 19801.Most frequent in <strong>the</strong> Newbridge-Pippingford area, and a few plants near Old Lodge3 -++---"""''"---'--1- and in <strong>the</strong> wet woodlands by Pooh Bridge. The extensive stands at Newbridge Bog in<strong>the</strong> alder coppice grow in a thick iron 'soup', Where <strong>the</strong> coppice is very dense <strong>the</strong> plantscan no longer stand <strong>the</strong> shade, stop flowering and produce few and fewer leaves until<strong>the</strong> tussocks die; <strong>the</strong> dead tussocks remain under <strong>the</strong> alders for some years like little4 5gravestones. However, after coppicing <strong>of</strong> <strong>the</strong> alders in 1994, seed germinated inabundance in <strong>the</strong> open areas, and plants flowered from <strong>the</strong>ir second year. The development <strong>of</strong> <strong>the</strong> inflorescence atNewbridge is very variable; inflorescences may be dense or lax in <strong>the</strong> open and sparse in shade, branched qr simple,usually without but sometimes with bracts, and <strong>the</strong> fruits can be broad or quite narrow.Very locally frequent in Sussex in <strong>the</strong> Weald. Widespread in lowland Britain <strong>of</strong>ten in peaty, medium-rich,seasonally wet soils. Fairly widespread in Europe, Caucasus and western North Africa.Carex x boenninghausiana (C. paniculata x remota).Newbridge Bog, P. C. Hall (Hall 19801. Searched for in 1994 and not found but as many<strong>of</strong> <strong>the</strong> big Carex tussocks were not flowering in <strong>the</strong> woodland due to dense shade itwould have been very difficult to spot; it could still survive.One large plant was found in amongst C. remota in an alder flush in Pippingford3 Park just south-east <strong>of</strong> <strong>the</strong> Centre Bridge (451.3161, 1995, SBRS/Kent Field Clubmeeting, although <strong>the</strong>re was no C. paniculata anywhere near (confirmed A. O. Chater);<strong>the</strong> plant looks like C. paniculata but has narrower leaves and long bracts.4 5
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Flora of Ashdown Forest,-,;I
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Copyright Tim Rich, Pat Donovan, Pa
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ACKNOWLEDGEMENTSWe would like to th
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THE HABITATS OF ASH DOWN FOREST3His
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Near the bottom of all the non~wood
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Conopodium majusCrataegus laevigata
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Fincham (1995) published a small ph
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13ANAL VS IS OF THE FLORA DATAIn th
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15The number of different visits to
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17Hedgerows Disturbed/waste ground
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19The frequencies with which specie
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21(r~0.434, pO.l).-0.059,•300t350
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23A second type of comparison has b
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25Similarly, environmental variable
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SPECIES ACCOUNTS27Lat'1n names foll
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29EQUISETACEAEEquisetum f/uviatile.
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One plant on a vertical, north-faci
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Viola pa/ustris in an open alder co
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35DRYOPTERIDACEAEPolystichum aculea
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37BLECHNACEAEBlechnum spicant. Hard
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39AngiospermsLAURACEAE* Laurus nobi
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41Ranunculus repens, Creeping butte
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wet mud, small streams, ditches and
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45*Meconopsis cambrica. Welsh poppy
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47Ulmusx hollandica f?U. glabra x m
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49FAGACEAEFagus sylvatica. Beech.Fo
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A special large oak at Chuck Hatch,
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Cory/us avel/ana. Hazel.Recorded in
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plants in the autumn; we have not r
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Sagina procumbens. Procumbent pearl
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59*Saponaria officinalis. Soapwort,
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61Dolygonum arenastrum. Equal-leave
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63Rumex conglomeratus. Clustered do
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65Hypericum humifusum. Trailing S1
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67DROSERACEAEDrosera rotundifolia.
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69Viola laetea. Pale dog-violet.Thi
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71* Salix alba. White willow.3 -+-t
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73Salix repens var. repens (var. er
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75*Rorippa sylvestris. Creeping yel
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77* Thlaspi arvense. Field pennycre
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79aphanus raphanistrum. Wild radish
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ica tetralix. Cross~leaved heath.As
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centre of the Forest on alluvial so
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It is predominantly a plant which o
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87SAXIFRAGACEAE[*Darmera pe/tata. I
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89insipid in flavour IHaskel1 1960)
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91ection Rubus. Subsection Hiemales
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93Rubus pr%ngatus.Nutley (45.27), 1
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95bus vestitus.Newbridge, 1969, B.
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97Rubus largificus.3 -+t--------'+_
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Potentilla anserina. Silverweed.Com
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101Geum urbanum. Wood avens.Primari
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103osa x pseudorusticana fR. arvens
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105Prunus spinosa. Blackthorn, Wint
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orbus aucuparia. Rowan, Mountain as
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109r*Crataegus persimilis. Broad-le
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111icia tetrasperma. Smooth tare.Fo
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113Trifolium rep ens. White clover"
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115Genista tinctoria. Dyer's greenw
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*Genista anglica. Petty whin, Needl
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Myriophyllum alterniflorum. Alterna
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121Epilobium montanum x obscurum.Fo
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habitats.It lives up to the name of
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125,UPHORBIACEAEVlercurialis perenn
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Although one of Britain's tiniest p
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129*Oxalis articulata. Pink-sorrel.
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*Erodium cicutarium. Common stork's
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Widespread in Europe from the shore
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* Sison amomum. Stone parsley.Tetra
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137Centaurium pu/chellum. Lesser ce
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139CONVOLVULACEAEConvolvulus arvens
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141*Nymphoides pe/tata. Fringed wat
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Myosotis secunda (M. repens). Creep
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145Stachys 8lvensis. Field woundwor
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5cutellaria x hybrida (5. galericul
- Page 154 and 155: Origanum vu/gare. Marjoram.Near Fis
- Page 156 and 157: CALLlTRICHACEAE151Ca/litriche is a
- Page 158 and 159: 153Fraxinus excelsior. Ash.4 5Local
- Page 160 and 161: ~'Cymbalaria mural is. Ivy-leaved t
- Page 162 and 163: 157Veronica scutellata. Marsh speed
- Page 164 and 165: 159Sibthorpia europaea. Cornish mon
- Page 166 and 167: 161OROBANCHACEAEOrobanche rapum~gen
- Page 168 and 169: 163RUBIACEAESherardia arvensis. Mad
- Page 170 and 171: 165Viburnum OpUIU5.Guelder-rose, St
- Page 172 and 173: 167Valeriana officina/is 5.1. Commo
- Page 174 and 175: 169Cirsium vu/gate. Spear thistle.W
- Page 176 and 177: 171We suspect that Leontodon hispid
- Page 178 and 179: 173Mycelis muralis. Wall lettuce.Se
- Page 180 and 181: 175Taraxacum exceJlens.Scattered al
- Page 182 and 183: 177Taraxacum sinuatum.Recorded from
- Page 184 and 185: 179Crepis capillaris. Smooth hawk'
- Page 186 and 187: 181Hieracium. Hawkweeds.3134 5Hiera
- Page 188 and 189: *So/idago gigantea. Early goldenrod
- Page 190 and 191: ecorded with Chamaemelum are Matric
- Page 192 and 193: *Senecio x albescens (S. cineraria
- Page 194 and 195: *Petasites japonicus. Giant butterb
- Page 196 and 197: *E1odea canadensis. Canadian waterw
- Page 198 and 199: Arum maculatum. Lords~and~ladies, C
- Page 200 and 201: Juncus x surrejanus (J. acutiflorus
- Page 202 and 203: fences or topography. It may domina
- Page 206 and 207: A hybrid first described by Col em
- Page 208 and 209: 203Carex riparia. Greater pond~sedg
- Page 210 and 211: Material which initially looked lik
- Page 212 and 213: 207Festuca pratensis. Meadow fescue
- Page 214 and 215: 209Cynosurus cristatus. Crested dog
- Page 216 and 217: 211Glyceria f/uitans. Floating swee
- Page 218 and 219: 213Deschampsia cespitosa. Tufted ha
- Page 220 and 221: Agrostis gigantea. Black bent.Tetra
- Page 222 and 223: 217Bromus lepidus. Slender soft~bro
- Page 224 and 225: * Triticum aestivum. Bread wheat.A
- Page 226 and 227: 221L1L1ACEAENarthecium ossifragum.
- Page 228 and 229: 223*Allium triquetrum. Three-Gornar
- Page 230 and 231: Trumpet length(mm)454035302520•
- Page 232 and 233: 227ORCHIDACEAEEpipactis purpurata.
- Page 234 and 235: een unable to refind any bog orchid
- Page 236 and 237: 231DactylorhizB praetermissa. South
- Page 238 and 239: Rachel A. Nicholson (1934-). Rachel
- Page 240 and 241: 235REFERENCESAbbas, M. H. (1979). S
- Page 242 and 243: Graham, G. G. & Primavesi, A. L. (1
- Page 244 and 245: Rich, T. C. G. & Smith, P. A. (1996
- Page 246 and 247: GAZETTEER241This gazetteer has been
- Page 248 and 249: Smockfarthing 40.31Smuggler's car p
- Page 250 and 251: Asple adi+cuttr int~~~tl·Bteel SI!
- Page 252 and 253: Broad bean 111 Cardamine x hausskne
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Fen bedstraw 163 Fumitory 46 Greate
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Leucanthemum maximum 186 Many-flowe
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Quercus ilex 50 Rorippa x ster/lis
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Sticky mouse-ear 56 Thale cress 73