Arum maculatum. Lords~and~ladies, Cuckoo pint, Lily grass, Small dragon, Great dragon, Ram's horn, Wake Robin,Schoolmaster.Recorded widely around <strong>the</strong> <strong>Forest</strong> in hedges, woodland, gardens and road verges,usually on <strong>the</strong> richer soils,Recorded in 92% <strong>of</strong> <strong>the</strong> tetrads in Sussex (Hall 1980). Widespread in Britain andreaching its nor<strong>the</strong>rn limit in sou<strong>the</strong>rn Scotland. Western, central and sou<strong>the</strong>rn Europe,and North Africa.1934*Arum italicum subsp. italicum. Italian lords-and-Iadies.Several clumps established in lane east <strong>of</strong> ford at Oldlands Farm {476.2681, leaping <strong>of</strong>fdown lane from <strong>the</strong> farm garden, 1995, TR & PA; one plant in hedge by Duddleswellcross-roads {471 .2881. 1995, RN & ER.Subsp. italicum is increasing as a garden escape in England (Rich & Woodruff3 -++------Lj- 1996), and is native in sou<strong>the</strong>rn and western Europe and North Africa. None <strong>of</strong> <strong>the</strong>native 5ubsp. neglectum has been found, and would probably not be expected (Prime,Buckle & Lovis 1955).4 5LEMNACEAE* Spirode/a po/yrhiza. Greater duckweed.A few plants in a new pond in Pippingford Park {448.2991, 1995, TR, where it wasprobably an accidental introduction w'lth o<strong>the</strong>r aquatics,Locally abundant in rivers, ditches and ponds in Sussex, especially on <strong>the</strong> grazingmarshes <strong>of</strong> areas such as Pevensey and Lewes, and locally distributed in lowland Britain,3 Widespread in Europe except <strong>the</strong> extreme north and south, and many o<strong>the</strong>r places in <strong>the</strong>world.4 5Lemna minor. Common duckweed.Common on <strong>the</strong> <strong>Forest</strong> {Hall 19801.Scattered mainly around <strong>the</strong> edges <strong>of</strong> <strong>the</strong> <strong>Forest</strong>, in small flushes, open and shadedponds, and in <strong>the</strong> Medway. This is our commonest duckweed, as it is in Britain.It is widespread in Sussex in a wide range <strong>of</strong> aquatic habitats and a range <strong>of</strong> water3 -++---~--'+- conditions. L minor is able to grow optimally at lower levels <strong>of</strong> phosphate in <strong>the</strong> waterthan <strong>the</strong> o<strong>the</strong>r duck weeds, whilst L minuta is able to grow at lower nitrogenconcentrations (Lound 1980), It occurs throughout lowland Britain and Europe,4 5Lemna trisu/ca, Ivy-leaved duckweed.,----,-==---'--,-----, Tetrad 43 B (H a 11 1980).Pond at Ridge Road (446.328), 1993, TR, but not seen in 1995.This submerged duckweed is quite shade-tolerant, <strong>of</strong>ten growing under o<strong>the</strong>rfloating aquatics and may- not be noticed unless fished out with a grapnel or stick, It is3 -++---------'_+_locally distributed in Sussex, and widespread in lowland Britain and Europe, It iscosmopolitan except for <strong>the</strong> polar regions and <strong>the</strong> tropics,4 5
194*Lemna minuta (L. minuscula), Least duckweed.Pond south <strong>of</strong> Nutley 1447.2631, 1993, MM; Marlpits pond (467.263), 1993, TR & NM;pond in Furnace Wood (476.2601, 1995, TR; pulled up on grapnel with Callitriche fromRiver Medway (443.3491, 1995, AK & TR, but only noticed when we tried to identify <strong>the</strong>Callitriche at home!3 ++-------L.f- This species was first recognised in Britain in 1977 (Leslie & Waiters 1983), and hasdefinitely been spreading ever since and was not just overlooked. It was not recorded inHall 11980) and only a few times in Briggs (19901, but has continued to spread since. Itis now widespread in sou<strong>the</strong>rn Britain. Native in North and South America.4 5JUNCACEAEJunCU5 squarrosus. Heath rush, Goose corn.On <strong>Ashdown</strong> <strong>Forest</strong>, not common IColeman 1836). Near Pipping ford, 1904, J. Stirling(TLS). Thinly scattered on <strong>Ashdown</strong>, near Gills Lap, etc. (Done 1914). <strong>Forest</strong> nearPippingford, J. Stirling; <strong>Ashdown</strong> <strong>Forest</strong>, locally common, A. H. Wolley-Dod (Wolley-Dod1937). Common on <strong>the</strong> <strong>Forest</strong> (Hall 1980).Widely distributed on rides, acidic grassland and wet patches on heath land, tolerant<strong>of</strong> grazing and trampling. It is a very characteristic heathland species.This species keeps its place in <strong>the</strong> vegetation by forcing its leaves outwards anddown, pushing its competitors out <strong>of</strong> <strong>the</strong> way. The patches slowly increase in size andeventually form rings as <strong>the</strong>y decay in <strong>the</strong> centre. At King's Standing it is frequent intrampled areas near <strong>the</strong> car park, forming small patches and decaying rings mostly up to 20 cm diameter.Widespread in nor<strong>the</strong>rn and western Britain, and on <strong>the</strong> heaths in East Anglia, <strong>the</strong> Weald, Surrey and <strong>the</strong> New<strong>Forest</strong>. Central, western and north-west Europe, Greenland and North Africa.*Juncus tenuis (J. macer). Slender rush.Newbridge, near Coleman's Hatch, Miss K. Pickard; near Old Mill, Crowborough Warren,T. Hilton; near Maynard's Gate, A. H. WoIJey-Dod; in two spots near Wych Cross, A. H.Wolley-Dod (Wolley-Dod 1937), and described as rare but spreading. Abundant on paths,Hindleap Warren and Broadstone Warren, 1948, R. A. Boniface. Frequent on <strong>the</strong> <strong>Forest</strong>(Hall 1980).Widely distributed on tracks, around car parks and especially on rides in woodlands,<strong>of</strong>ten in damp, shaded situations but also sometimes in dry, open places. Its spreadseems to be especially associated with forestry operations.Becoming widespread in Britain, especially in <strong>the</strong> north and west, and naturalized innor<strong>the</strong>rn, western and central Europe. It is native in North and South America.Juncus bufonius sensu stricto. Toad rush.Common on <strong>the</strong> <strong>Forest</strong> (Hall 1980).Common everywhere on rides, pond edges, damp patches, disturbed clayey ground,car parks and sometimes road gutters.It mainly grows on open mud <strong>of</strong> pH 4-5, but will grow on o<strong>the</strong>r soils too, with an3estimated seed output <strong>of</strong> 13-15 million seeds per square metre (Salisbury 1970). It isone <strong>of</strong> <strong>the</strong> most widespread mud species, almost cosmopolitan, and increasing in England(Rich & Woodruff 1996).4 5Juncus articulatus (J. lampocarpus). Jointed rush.<strong>Ashdown</strong> <strong>Forest</strong> (Done 1914). Common on <strong>the</strong> <strong>Forest</strong> (Hall 1980).Widespread but not common on <strong>the</strong> <strong>Forest</strong>, usually on open sides <strong>of</strong> streams andponds where <strong>the</strong> vegetation is slightly open, sometimes on woodland rides in unshadedplaces.Widespread in Britain and Europe, and in <strong>the</strong> nor<strong>the</strong>rn hemisphere.This species and J. acutiflorus can be difficult to distinguish until in fruit (Rich &Rich 1988). J. articulatus has blackish fruits with a distinct point (acumen) at <strong>the</strong> tip. J.acutiflorus has brownish fruits which taper to <strong>the</strong> tip.
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Flora of Ashdown Forest,-,;I
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Copyright Tim Rich, Pat Donovan, Pa
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ACKNOWLEDGEMENTSWe would like to th
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THE HABITATS OF ASH DOWN FOREST3His
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Near the bottom of all the non~wood
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Conopodium majusCrataegus laevigata
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Fincham (1995) published a small ph
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13ANAL VS IS OF THE FLORA DATAIn th
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15The number of different visits to
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17Hedgerows Disturbed/waste ground
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19The frequencies with which specie
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21(r~0.434, pO.l).-0.059,•300t350
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23A second type of comparison has b
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25Similarly, environmental variable
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SPECIES ACCOUNTS27Lat'1n names foll
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29EQUISETACEAEEquisetum f/uviatile.
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One plant on a vertical, north-faci
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Viola pa/ustris in an open alder co
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35DRYOPTERIDACEAEPolystichum aculea
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37BLECHNACEAEBlechnum spicant. Hard
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39AngiospermsLAURACEAE* Laurus nobi
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41Ranunculus repens, Creeping butte
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wet mud, small streams, ditches and
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45*Meconopsis cambrica. Welsh poppy
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47Ulmusx hollandica f?U. glabra x m
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49FAGACEAEFagus sylvatica. Beech.Fo
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A special large oak at Chuck Hatch,
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Cory/us avel/ana. Hazel.Recorded in
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plants in the autumn; we have not r
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Sagina procumbens. Procumbent pearl
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59*Saponaria officinalis. Soapwort,
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61Dolygonum arenastrum. Equal-leave
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63Rumex conglomeratus. Clustered do
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65Hypericum humifusum. Trailing S1
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67DROSERACEAEDrosera rotundifolia.
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69Viola laetea. Pale dog-violet.Thi
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71* Salix alba. White willow.3 -+-t
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73Salix repens var. repens (var. er
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75*Rorippa sylvestris. Creeping yel
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77* Thlaspi arvense. Field pennycre
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79aphanus raphanistrum. Wild radish
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ica tetralix. Cross~leaved heath.As
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centre of the Forest on alluvial so
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It is predominantly a plant which o
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87SAXIFRAGACEAE[*Darmera pe/tata. I
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89insipid in flavour IHaskel1 1960)
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91ection Rubus. Subsection Hiemales
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93Rubus pr%ngatus.Nutley (45.27), 1
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95bus vestitus.Newbridge, 1969, B.
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97Rubus largificus.3 -+t--------'+_
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Potentilla anserina. Silverweed.Com
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101Geum urbanum. Wood avens.Primari
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103osa x pseudorusticana fR. arvens
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105Prunus spinosa. Blackthorn, Wint
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orbus aucuparia. Rowan, Mountain as
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109r*Crataegus persimilis. Broad-le
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111icia tetrasperma. Smooth tare.Fo
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113Trifolium rep ens. White clover"
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115Genista tinctoria. Dyer's greenw
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*Genista anglica. Petty whin, Needl
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Myriophyllum alterniflorum. Alterna
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121Epilobium montanum x obscurum.Fo
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habitats.It lives up to the name of
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125,UPHORBIACEAEVlercurialis perenn
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Although one of Britain's tiniest p
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129*Oxalis articulata. Pink-sorrel.
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*Erodium cicutarium. Common stork's
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Widespread in Europe from the shore
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* Sison amomum. Stone parsley.Tetra
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137Centaurium pu/chellum. Lesser ce
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139CONVOLVULACEAEConvolvulus arvens
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141*Nymphoides pe/tata. Fringed wat
- Page 148 and 149: Myosotis secunda (M. repens). Creep
- Page 150 and 151: 145Stachys 8lvensis. Field woundwor
- Page 152 and 153: 5cutellaria x hybrida (5. galericul
- Page 154 and 155: Origanum vu/gare. Marjoram.Near Fis
- Page 156 and 157: CALLlTRICHACEAE151Ca/litriche is a
- Page 158 and 159: 153Fraxinus excelsior. Ash.4 5Local
- Page 160 and 161: ~'Cymbalaria mural is. Ivy-leaved t
- Page 162 and 163: 157Veronica scutellata. Marsh speed
- Page 164 and 165: 159Sibthorpia europaea. Cornish mon
- Page 166 and 167: 161OROBANCHACEAEOrobanche rapum~gen
- Page 168 and 169: 163RUBIACEAESherardia arvensis. Mad
- Page 170 and 171: 165Viburnum OpUIU5.Guelder-rose, St
- Page 172 and 173: 167Valeriana officina/is 5.1. Commo
- Page 174 and 175: 169Cirsium vu/gate. Spear thistle.W
- Page 176 and 177: 171We suspect that Leontodon hispid
- Page 178 and 179: 173Mycelis muralis. Wall lettuce.Se
- Page 180 and 181: 175Taraxacum exceJlens.Scattered al
- Page 182 and 183: 177Taraxacum sinuatum.Recorded from
- Page 184 and 185: 179Crepis capillaris. Smooth hawk'
- Page 186 and 187: 181Hieracium. Hawkweeds.3134 5Hiera
- Page 188 and 189: *So/idago gigantea. Early goldenrod
- Page 190 and 191: ecorded with Chamaemelum are Matric
- Page 192 and 193: *Senecio x albescens (S. cineraria
- Page 194 and 195: *Petasites japonicus. Giant butterb
- Page 196 and 197: *E1odea canadensis. Canadian waterw
- Page 200 and 201: Juncus x surrejanus (J. acutiflorus
- Page 202 and 203: fences or topography. It may domina
- Page 204 and 205: Eleocharis multicaulis. Many-stalke
- Page 206 and 207: A hybrid first described by Col em
- Page 208 and 209: 203Carex riparia. Greater pond~sedg
- Page 210 and 211: Material which initially looked lik
- Page 212 and 213: 207Festuca pratensis. Meadow fescue
- Page 214 and 215: 209Cynosurus cristatus. Crested dog
- Page 216 and 217: 211Glyceria f/uitans. Floating swee
- Page 218 and 219: 213Deschampsia cespitosa. Tufted ha
- Page 220 and 221: Agrostis gigantea. Black bent.Tetra
- Page 222 and 223: 217Bromus lepidus. Slender soft~bro
- Page 224 and 225: * Triticum aestivum. Bread wheat.A
- Page 226 and 227: 221L1L1ACEAENarthecium ossifragum.
- Page 228 and 229: 223*Allium triquetrum. Three-Gornar
- Page 230 and 231: Trumpet length(mm)454035302520•
- Page 232 and 233: 227ORCHIDACEAEEpipactis purpurata.
- Page 234 and 235: een unable to refind any bog orchid
- Page 236 and 237: 231DactylorhizB praetermissa. South
- Page 238 and 239: Rachel A. Nicholson (1934-). Rachel
- Page 240 and 241: 235REFERENCESAbbas, M. H. (1979). S
- Page 242 and 243: Graham, G. G. & Primavesi, A. L. (1
- Page 244 and 245: Rich, T. C. G. & Smith, P. A. (1996
- Page 246 and 247: GAZETTEER241This gazetteer has been
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Smockfarthing 40.31Smuggler's car p
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Asple adi+cuttr int~~~tl·Bteel SI!
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Broad bean 111 Cardamine x hausskne
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Fen bedstraw 163 Fumitory 46 Greate
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Leucanthemum maximum 186 Many-flowe
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Quercus ilex 50 Rorippa x ster/lis
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Sticky mouse-ear 56 Thale cress 73