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Flora of Ashdown Forest - Botanical Society of the British Isles

Flora of Ashdown Forest - Botanical Society of the British Isles

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60Persicaria amphibia (Po/ygonum amphibiumj. Amphibious bistort.Tetrads 43A and 43H IHall 19801.Surprisingly rare given <strong>the</strong> number <strong>of</strong> ponds with base-poor, medium-nutrient water, andonly recorded in 47.26, 1994, <strong>Flora</strong> meeting Idetails not recollected and possibly an error),One <strong>of</strong> <strong>the</strong> few truly amphibious plants in Britain, occurring in permanent water and on3 -++------'-+_ dry land though <strong>of</strong>ten not flowering in <strong>the</strong> latter case. Common in Britain and increasing (Rich& Woodruff 1996), and widespread in Europe, Asia, Africa and North America.4 5Persicaria maculosa (Polygonum persicariaj. Redshank, Lover's pride.Frequent on roads ides especially in <strong>the</strong> gutters, on pond edges, in <strong>the</strong> villages, car parks andarable fields, etc., <strong>of</strong>ten on reasonably nutrient-rich soils. It is only occasionally found inwoodland rides and does not appear as shade-tolerant as P. hydropiper. Whilst most plantsexamined have lacked glands, a few with glands scattered on <strong>the</strong> pedicels were treated asthis species and not <strong>the</strong> next.Recorded in 90% <strong>of</strong> <strong>the</strong> tetrads in Sussex (Hall 19801. Common throughout lowlandBritain and Europe, and widespread elsewhere in <strong>the</strong> world as a weed.Persicaria lapathifolia (Polygonum lapathifolium). Pale persicaria.Common on <strong>the</strong> <strong>Forest</strong> IHall 19801.Possibly under-recorded on <strong>the</strong> <strong>Forest</strong>. This and P. maculosa frequently grow toge<strong>the</strong>rbut <strong>the</strong> differences are not always immediately obvious; P. lapathifolia is certainly lesscommon. It is found on <strong>the</strong> edges <strong>of</strong> roads, in gardens and in arable fields. It was abundant3 --+.--------'_+_ at <strong>the</strong> organic farm at Plawhatch in 1995.Common in Sussex and lowland Britain, and possibly increasing in England (Rich &Woodruff 1996). Probably more widespread in Europe than P. maculosa. Found in temperateregions <strong>of</strong> <strong>the</strong> nor<strong>the</strong>rn hemisphere and in South Africa.4 5Persicaria hydropiper (Polygonum hydropiper). Water-pepper,Common on <strong>the</strong> <strong>Forest</strong> IHall 19801.Most typical <strong>of</strong> wet patches on shaded rides and <strong>of</strong>ten abundant on <strong>the</strong>m. It is especiallyfrequent in damp muddy hollows where <strong>the</strong> soils are silty and compa~ted, and it is largelyabsent from <strong>the</strong> heathy, acidic soils. It has a persistent seed bank and 4% <strong>of</strong> <strong>the</strong> seeds maystill be viable after 50 years ISalisbury 19701.Young plants can be distinguished from P. maculosa, with which it may grow, by <strong>the</strong>peppery taste as <strong>the</strong> English name hints.Common in Sussex, Britain, lowland Europe, temperate Asia snd North Africa.Persicaria minor (Polygonum minus). Small water-pepper.Near Rifle Butts, <strong>Forest</strong> Row, Miss P. Stockdale IBEX; Wolley-Dod 19371; this site was probably at about 425.337 andwas drained for <strong>the</strong> golf course.Probably extinct. This is a species <strong>of</strong> wet grassland, swamps and pond edges, <strong>of</strong>ten on nutrient-rich mud which driesout in summer (Mountford 1994). Plants produce both flat and trigonous seeds in varying proportions which differ in somegermination characteristics, though both germinate best in <strong>the</strong> spring and in <strong>the</strong> light (Salisbury 1970).Scattered thinly over <strong>the</strong> sou<strong>the</strong>rn part <strong>of</strong> Britain and always scarce in Sussex, particularly so in <strong>the</strong> east. Quitewidespread in Europe except <strong>the</strong> extreme north and south and temperate Asia.Polygonum aviculare sensu stricto. Knotgrass, Armstrong.Quite frequent in car parks, on road edges, field edges and villages, but very uncommonanywhere else.Virtually ubiquitous in lowland Europe, and temperate Asia.P. aviculare s.s. and P. arenastrum have <strong>of</strong>ten been recorded as an aggregate in <strong>the</strong> past,but <strong>the</strong>y are readily distinguished. P. aviculare is a sprawling plant with a variety <strong>of</strong> leaf sizes,and <strong>the</strong> fruit has three equal sides. P. arenastrum is a more compact plant with mainly equal,small leaves except at <strong>the</strong> tips <strong>of</strong> branches, and <strong>the</strong> fruits have two equal sides and a smaller,concave one.

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