109r*Crataegus persimilis. Broad-leaved cockspur-thorn.bout ten scattered trees planted in Raven Wood (431.311 I, 1995, TR. These trees are not regenerating at Raven Wood,ut are known to be bird-sown and established elsewhere in sou<strong>the</strong>rn England. They differ from C. crus-gall! in havinglightly hairy veins on <strong>the</strong> undersides <strong>of</strong> <strong>the</strong> leaves.]Crataegus monogyna. Hawthorn, May, Cuckoo's bread-and-cheese tree, Agarve, Agasse, Hogarve, Hoghazel, Hog-go5se.Recorded in every square on <strong>the</strong> <strong>Forest</strong>, in woods, hedges, scrub and on roadsides; rapidlycolonising drier neglected grassland sites on <strong>the</strong> better soils, and only rarely a primary colonist<strong>of</strong> heaths, for example near Gills Lap 1466.317). Cultivars with pink, double flowers areplanted around Nutley.This species tends to set fruit when cross-pollinated and may be largely selfincompatible.The fruits turned red at <strong>the</strong> beginning <strong>of</strong> September 1995, and were stillpresent on <strong>the</strong> bushes in February 1996 but <strong>the</strong>n disappeared rapidly in <strong>the</strong> cold wea<strong>the</strong>r.They are dispersed by resident and migratory birds and small mammals.Without doubt <strong>the</strong> commonest species planted in hedges in Sussex, <strong>of</strong>ten with Cory/usave//ana. The 'Isle <strong>of</strong> Thorns' is an enclosure made <strong>of</strong> thorns, and <strong>the</strong> hedge includes at least some hawthorn. The Sussexxpression 'ga<strong>the</strong>ring knots <strong>of</strong> may' has become corrupted by some to 'ga<strong>the</strong>ring nuts in May'.Ubiquitous in Sussex, lowland Britain and Europe east to Afghanistan.Crataegus laevigata x monogyna. Hybrid hawthorn.This hybrid is quite frequent around <strong>the</strong> <strong>Forest</strong> as scattered plants, and may occur in <strong>the</strong>absence <strong>of</strong> ei<strong>the</strong>r parent, but usually occurs near C. /aevigata. It tends to occur in open• woodland and in hedges, possibly where originally planted.Byatt (1975) analysed Crataegus populations in south-east England lincluding <strong>Ashdown</strong><strong>Forest</strong> - <strong>the</strong> grid reference should be TQ/42.30; pers. comm. 1996) and found that mostpopulations contain hybrids and show varied degrees <strong>of</strong> introgressive hybridisation.It is widely recorded in lowland Britain.Crataegus laevigata. Midland hawthorn.<strong>Forest</strong> Row, H. F. Parsons (Wolley-Dod 1937). Seven tetrad records including some in <strong>the</strong>middle <strong>of</strong> <strong>the</strong> <strong>Forest</strong> IHall 1980).Most typically a plant <strong>of</strong> woodland on heavy clay soils IByatt 1975) as shown clearly by<strong>the</strong> map, and usually regarded as an ancient woodland plant. Where it is frequent this may betrue, but it has recolonised a number <strong>of</strong> secondary woodland sites on <strong>the</strong> edge <strong>of</strong> <strong>the</strong> <strong>Forest</strong>,as at Priory Road 1412.338-1 and Toll Lane 1461.2631, presumably bird-sown. The mostunusual site was one bush on <strong>the</strong> north verge <strong>of</strong> Ridge Road c. 50 metres east <strong>of</strong> <strong>the</strong><strong>Ashdown</strong> Park Hotel entrance 1429.324).Crataegus /aevigata and <strong>the</strong> hybrid tend to flower a couple <strong>of</strong> weeks before C. monogynaand can <strong>the</strong>n be picked out for examination in more detail later.Widespread in <strong>the</strong> Weald, especially on <strong>the</strong> clays, and in lowland south-east Britain. It is endemic to Europe.FABACEAE*Robinia pseudacacia. False acacia.Two well-grown trees outside 'Dell Wood', Tompset's Bank 142.331, 1995, PW; <strong>Forest</strong> Row142.341, 1995, PO & DB; spreading out <strong>of</strong> gardens near Bank Cottage 143.331, 1995, TR;large tree outside Oldlands Farm 1477.268), 1995, TR.Often planted and occasionally naturalized (probably mostly spreading vegetatively).]--++-----'-+-Occasional in Sussex and increasingly recorded in England especially on sandy soils (Clement& Foster 1994). Native in North America.4 5
110Lotus corniculatus. Common bird's-foot-trefoil, Boots-and-shoes, Cuckoo's stockings, Eggs-and-bacon, Lady's fingers, MiI~maidens, Pattens-and-clogs, Pig's pettitoes, Shoes-and- stockings and many more!iWe have it from every square in car parks, along rides and in pastures. It seems tolerant 0\quite wet ground.iUbiquitous in Sussex (Hall 1980) and Britain, probably <strong>the</strong> most ecologically wide-ranginglegume in Britain. Widespread in Europe, Asia, North Africa and <strong>the</strong> East African mountains.Large continental forms <strong>of</strong> this plant are <strong>of</strong>ten sown on road verges in so-called ' wild,flower seed mixtures'. An upright form was noted on <strong>the</strong> verges <strong>of</strong> <strong>the</strong> A22 north <strong>of</strong> Nutley,(44.29), 1994, TR but its ongin is unknown.Lotus pedunculatus (L. uliginosus). Greater bird's-foot-trefoil.Common on <strong>the</strong> <strong>Forest</strong> (Hall 1980). Locally abundant on wet soils and in <strong>the</strong> flushes;sometimes forming quite large clumps, usually in ungrazed or lightly grazed sites.It has nitrogen-fixing nodules in <strong>the</strong> roots with <strong>the</strong> bacterium Rhizobium lupini, as havemost legumes. Grime et al. (1988) note that legumes may be rare in wetlands as <strong>the</strong>;nitrogen-fixing processes need oxygen which is in short supply in water-logged soils, and this:species is <strong>the</strong> only common wetland legume in Britain. It tends to occur in <strong>the</strong> drier parts <strong>of</strong>iwetlands, and it is fairly sensitive to ferrous iron toxicity.Widespread in Sussex and Britain. Widespread in west, central and sou<strong>the</strong>rn Europe,North Africa and <strong>the</strong> Canary Islands.Vicia cracca. Tufted vetch.Widespread but never common on <strong>the</strong> <strong>Forest</strong>, and usually on verges or climbing up hedges;<strong>the</strong> purple tufts <strong>of</strong> flowers are more readily spotted in hedges than <strong>the</strong> low-growing plants onroads ides which are <strong>of</strong>ten mown and do not flower. It needs o<strong>the</strong>r vegetation for support, andmanages to pull itself up by <strong>the</strong> tendrils at <strong>the</strong> ends <strong>of</strong> <strong>the</strong> leaves.Ubiquitous in Sussex (Hall 1980), and widespread in Britain. Increasing in England (Rich& Woodruff 1996). Widespread in Europe and Asia; in Japan and Greenland.Vicia hirsuta. Hairy tare.Records in Hall (1980) for <strong>the</strong> <strong>Flora</strong> area are confined to tetrad 42T (not refoundl, although itWe have found it in four sites: tens <strong>of</strong> plants with Filago minima on road verge near Goatcross-roads (397.332), 1992, AK and seen <strong>the</strong>re until 1994; lawns at Isle <strong>of</strong> Thorns(449.265), pH 5.3, 1994, TR; sandy field north <strong>of</strong> Old Lodge (458.303), c. 1990, OK but notThis plant clearly prefers sandy ra<strong>the</strong>r than clayey soil and is relatively common on <strong>the</strong> Lower Greensand in Westbeen seen recently near <strong>the</strong> coast in <strong>the</strong> far south-east and in Buckhurst Park. Nationally its distribution is concentrated in<strong>the</strong> south-east but it is less common in Sussex than in <strong>the</strong> adjoining counties, and is decreasing in England (Rich &Ornithopus perpusillus. Bird's-foot.is recorded for two fur<strong>the</strong>r tetrads in 42L and 42X just to <strong>the</strong> south <strong>of</strong> <strong>Ashdown</strong> <strong>Forest</strong>.3-+~--~~--~- (418.304), 1995, <strong>Flora</strong> meeting; abundant on <strong>the</strong> north side <strong>of</strong> a track to Hunter's Farm4 5refound in 1994/5, OK & TR.In <strong>the</strong> site near Goat cross-roads it appeared after clearance <strong>of</strong> a road verge, but by 1995had disappeared as a result <strong>of</strong> <strong>the</strong> area becoming very overgrown. In November 1995numerous seedlings were found at <strong>the</strong> Hunter's Farm locality on bare ground created by <strong>the</strong> summer drought.Sussex. In East Sussex it only occurs in scattered localities where, apart from <strong>the</strong> sites in <strong>Ashdown</strong> <strong>Forest</strong>, it has onlyWoodruff 1996). Mainly western in Europe.3-4~-------..-L+-4 5Seven tetrads on <strong>the</strong> <strong>Forest</strong> (Hall 1980).<strong>Forest</strong> Row, verge <strong>of</strong> old railway (436.347), 1995, PW; soil dumped on roadside oppositERadio Station (473.292), 1994, SBRS; verge with disturbed soil, Marden's Hill (499.325), p~7.6, 1993 but gone by 1994, TR.The white to very pale lilac flowers <strong>of</strong> this species contrast with <strong>the</strong> distinctly pale blUEflowers <strong>of</strong> V. tetrasperma. Both V. hirsuta and V. tetrasperma are distinctly rare on thE<strong>Forest</strong>, with <strong>the</strong> latter being relatively more frequent despite being reported from fewer tetrad~in Hall (1980).Common and widespread in Sussex, lowland Europe, western Asia and North Africa.
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Flora of Ashdown Forest,-,;I
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Copyright Tim Rich, Pat Donovan, Pa
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ACKNOWLEDGEMENTSWe would like to th
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THE HABITATS OF ASH DOWN FOREST3His
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Near the bottom of all the non~wood
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Conopodium majusCrataegus laevigata
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Fincham (1995) published a small ph
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13ANAL VS IS OF THE FLORA DATAIn th
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15The number of different visits to
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17Hedgerows Disturbed/waste ground
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19The frequencies with which specie
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21(r~0.434, pO.l).-0.059,•300t350
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23A second type of comparison has b
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25Similarly, environmental variable
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SPECIES ACCOUNTS27Lat'1n names foll
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29EQUISETACEAEEquisetum f/uviatile.
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One plant on a vertical, north-faci
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Viola pa/ustris in an open alder co
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35DRYOPTERIDACEAEPolystichum aculea
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37BLECHNACEAEBlechnum spicant. Hard
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39AngiospermsLAURACEAE* Laurus nobi
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41Ranunculus repens, Creeping butte
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wet mud, small streams, ditches and
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45*Meconopsis cambrica. Welsh poppy
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47Ulmusx hollandica f?U. glabra x m
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49FAGACEAEFagus sylvatica. Beech.Fo
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A special large oak at Chuck Hatch,
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Cory/us avel/ana. Hazel.Recorded in
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plants in the autumn; we have not r
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Sagina procumbens. Procumbent pearl
- Page 64 and 65: 59*Saponaria officinalis. Soapwort,
- Page 66 and 67: 61Dolygonum arenastrum. Equal-leave
- Page 68 and 69: 63Rumex conglomeratus. Clustered do
- Page 70 and 71: 65Hypericum humifusum. Trailing S1
- Page 72 and 73: 67DROSERACEAEDrosera rotundifolia.
- Page 74 and 75: 69Viola laetea. Pale dog-violet.Thi
- Page 76 and 77: 71* Salix alba. White willow.3 -+-t
- Page 78 and 79: 73Salix repens var. repens (var. er
- Page 80 and 81: 75*Rorippa sylvestris. Creeping yel
- Page 82 and 83: 77* Thlaspi arvense. Field pennycre
- Page 84 and 85: 79aphanus raphanistrum. Wild radish
- Page 86 and 87: ica tetralix. Cross~leaved heath.As
- Page 88 and 89: centre of the Forest on alluvial so
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- Page 92 and 93: 87SAXIFRAGACEAE[*Darmera pe/tata. I
- Page 94 and 95: 89insipid in flavour IHaskel1 1960)
- Page 96 and 97: 91ection Rubus. Subsection Hiemales
- Page 98 and 99: 93Rubus pr%ngatus.Nutley (45.27), 1
- Page 100 and 101: 95bus vestitus.Newbridge, 1969, B.
- Page 102 and 103: 97Rubus largificus.3 -+t--------'+_
- Page 104 and 105: Potentilla anserina. Silverweed.Com
- Page 106 and 107: 101Geum urbanum. Wood avens.Primari
- Page 108 and 109: 103osa x pseudorusticana fR. arvens
- Page 110 and 111: 105Prunus spinosa. Blackthorn, Wint
- Page 112 and 113: orbus aucuparia. Rowan, Mountain as
- Page 116 and 117: 111icia tetrasperma. Smooth tare.Fo
- Page 118 and 119: 113Trifolium rep ens. White clover"
- Page 120 and 121: 115Genista tinctoria. Dyer's greenw
- Page 122 and 123: *Genista anglica. Petty whin, Needl
- Page 124 and 125: Myriophyllum alterniflorum. Alterna
- Page 126 and 127: 121Epilobium montanum x obscurum.Fo
- Page 128 and 129: habitats.It lives up to the name of
- Page 130 and 131: 125,UPHORBIACEAEVlercurialis perenn
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- Page 134 and 135: 129*Oxalis articulata. Pink-sorrel.
- Page 136 and 137: *Erodium cicutarium. Common stork's
- Page 138 and 139: Widespread in Europe from the shore
- Page 140 and 141: * Sison amomum. Stone parsley.Tetra
- Page 142 and 143: 137Centaurium pu/chellum. Lesser ce
- Page 144 and 145: 139CONVOLVULACEAEConvolvulus arvens
- Page 146 and 147: 141*Nymphoides pe/tata. Fringed wat
- Page 148 and 149: Myosotis secunda (M. repens). Creep
- Page 150 and 151: 145Stachys 8lvensis. Field woundwor
- Page 152 and 153: 5cutellaria x hybrida (5. galericul
- Page 154 and 155: Origanum vu/gare. Marjoram.Near Fis
- Page 156 and 157: CALLlTRICHACEAE151Ca/litriche is a
- Page 158 and 159: 153Fraxinus excelsior. Ash.4 5Local
- Page 160 and 161: ~'Cymbalaria mural is. Ivy-leaved t
- Page 162 and 163: 157Veronica scutellata. Marsh speed
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159Sibthorpia europaea. Cornish mon
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161OROBANCHACEAEOrobanche rapum~gen
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163RUBIACEAESherardia arvensis. Mad
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165Viburnum OpUIU5.Guelder-rose, St
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167Valeriana officina/is 5.1. Commo
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169Cirsium vu/gate. Spear thistle.W
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171We suspect that Leontodon hispid
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173Mycelis muralis. Wall lettuce.Se
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175Taraxacum exceJlens.Scattered al
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177Taraxacum sinuatum.Recorded from
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179Crepis capillaris. Smooth hawk'
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181Hieracium. Hawkweeds.3134 5Hiera
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*So/idago gigantea. Early goldenrod
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ecorded with Chamaemelum are Matric
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*Senecio x albescens (S. cineraria
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*Petasites japonicus. Giant butterb
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*E1odea canadensis. Canadian waterw
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Arum maculatum. Lords~and~ladies, C
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Juncus x surrejanus (J. acutiflorus
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fences or topography. It may domina
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Eleocharis multicaulis. Many-stalke
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A hybrid first described by Col em
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203Carex riparia. Greater pond~sedg
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Material which initially looked lik
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207Festuca pratensis. Meadow fescue
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209Cynosurus cristatus. Crested dog
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211Glyceria f/uitans. Floating swee
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213Deschampsia cespitosa. Tufted ha
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Agrostis gigantea. Black bent.Tetra
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217Bromus lepidus. Slender soft~bro
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* Triticum aestivum. Bread wheat.A
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221L1L1ACEAENarthecium ossifragum.
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223*Allium triquetrum. Three-Gornar
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Trumpet length(mm)454035302520•
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227ORCHIDACEAEEpipactis purpurata.
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een unable to refind any bog orchid
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231DactylorhizB praetermissa. South
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Rachel A. Nicholson (1934-). Rachel
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235REFERENCESAbbas, M. H. (1979). S
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Graham, G. G. & Primavesi, A. L. (1
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Rich, T. C. G. & Smith, P. A. (1996
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GAZETTEER241This gazetteer has been
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Smockfarthing 40.31Smuggler's car p
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Asple adi+cuttr int~~~tl·Bteel SI!
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Broad bean 111 Cardamine x hausskne
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Fen bedstraw 163 Fumitory 46 Greate
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Leucanthemum maximum 186 Many-flowe
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Quercus ilex 50 Rorippa x ster/lis
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Sticky mouse-ear 56 Thale cress 73