235REFERENCESAbbas, M. H. (1979). Soil formation and classification in <strong>the</strong> High Weald <strong>of</strong> Kent and Sussex with particular reference to<strong>Ashdown</strong> <strong>Forest</strong> and <strong>the</strong> development <strong>of</strong> compact subsoils. PhD Thesis, Wye College, University <strong>of</strong> London.Alien, D. E. 11954}. Variation in Peplis portula L. Watsonia 3: 85-91.Arnold, F. H. 11887}. <strong>Flora</strong> <strong>of</strong> Sussex. Simpkin, Marshall, Hamilton, Kent & Co., London.Arnold, F. H. 11907}. <strong>Flora</strong> <strong>of</strong> Sussex. 2nd edition. Simpkin, Marshall, Hamilton, Kent & Co., London.Arthur, D., ed. 11989}. A Sussex life. Barrie & Jenkins, London.Atkinson, M. D. 11992}. Biological <strong>Flora</strong> <strong>of</strong> <strong>the</strong> <strong>British</strong> <strong>Isles</strong>. Betula pendula Roth lB. verrucosa Ehrh.} and B. pubescensEhrh. Journal <strong>of</strong> Ecology 80: 837-870.Baker, H. G. 11947}. Biological <strong>Flora</strong> 01 <strong>the</strong> <strong>British</strong> <strong>Isles</strong>. Melandrium (Roehling) em. Fries genus. Journal <strong>of</strong> Ecology 35:271-292.Bannister, P. 11965}. Biological <strong>Flora</strong> <strong>of</strong> <strong>the</strong> <strong>British</strong> <strong>Isles</strong>. Erica cinerea L. Journal <strong>of</strong> Ecology 53: 527-542.Barkham, J. P. (1980a). Population dynamics <strong>of</strong> <strong>the</strong> wild daffodil (Narcissus pseudonarcissus). l. Clonal growth, seedreproduction, mortality and <strong>the</strong> effects <strong>of</strong> density. Journal <strong>of</strong> Ecology 68: 607-633.Barkham, J. P. (198Gb). Population dynamics <strong>of</strong> <strong>the</strong> wild daffodil (Narcissus pseudonarcissus). 11. Changes in <strong>the</strong> number<strong>of</strong> shoots and flowers, and <strong>the</strong> effect <strong>of</strong> bulb depth on growth and reproduction. Journal <strong>of</strong> Ecology 68: 635-664 ..Barling, D. H. (1958). Reproduction and cytology 01 Allium paradoxum. Proceedings BSBI3: 85.Barton, W. C. & Riddelsdell, H. J. 11936}. A new Rubus from Sussex. Journal <strong>of</strong> Botany 74: 204-205.Baskin, J. M. & Baskin, C. C. 11983}. Germination ecology <strong>of</strong> Veronica arvensis. Journal <strong>of</strong> Ecology 71: 57-68.Bates, H. E. 11943}. Country life. Penguin Books, London.Beattie, A. J. (1969). The floral biology <strong>of</strong> three species 01 Viola. New Phytologist 68: 1187-1201.Beattie, A. J. & Lyons, N. (1975). Seed dispersal in Viola (Violaceae): adaptations and strategies. American Journal <strong>of</strong>Botany 62: 714-722.Bevan, D. & Rich, T. C. G. (1991). Bevans' Bittercress: The white-flowered form <strong>of</strong> Cardamine X frings/i' Wirtgen presentin Britain. Watsonia 18: 403-405.Birkinshaw, C. R. (1994). Aspects <strong>of</strong> <strong>the</strong> ecology <strong>of</strong> Damasonium alisma Miller in Western Europe. Watsonia 20: 33-39.Bjorkman. O. & Powles, S. B. (1981). Leaf movement in <strong>the</strong> shade species Oxalis oregana. 1. Response to light level andquality. Annual Report <strong>of</strong> <strong>the</strong> Carnegie Institute, Washington 1980-1981: 59-62.Borrill, M. (1958a). A biosystematic study <strong>of</strong> some Glyceria species in Britain. 3. Biometrical studies. Watsonia 4: 77-88.Borrill, M. (1958b). A biosystematic study <strong>of</strong> some Glyceria species in Britain. 4. Breeding systems, fertility relationshipsand general discussion. Watsonia 4: 89-100.Boutin, C. & Harper, J. L. (1991), A comparative study <strong>of</strong> <strong>the</strong> population dynamics <strong>of</strong> five species <strong>of</strong> Veronica in naluralhabitats. Journal <strong>of</strong> Ecology 79: 199-221.Boyd, M., Silvertown, J. & Tucker, C. 11990}. Population ecology 01 heterostyle and homostyle Primula vUlgans: growth,survival and reproduction in field populations. Journal <strong>of</strong> Ecology 78: 799-813.Brewis, A., Bowman, P. & Rose, F. (1996). The flora <strong>of</strong> Hampshire. Harley Books, Colchester.Briggs, M. (1990), Sussex plant atlas. Selected supplement. Booth Museum, Brighton.Bristow, C. R. & Bazley, R. A. (1972), Geology <strong>of</strong> <strong>the</strong> country around Royal Tunbridge Wells. Memoirs <strong>of</strong> <strong>the</strong> GeologicalSurvey <strong>of</strong> Great Britain. HMSO, London.Byatt, J. I. (1975). Hybridisation between Crataegus manogyna Jacq. and C, laevigata (Poiret) DC. in South-easternEngland. Watsonia 10: 253-264.Bylield, A. J. 11994}. Cicendia filiformis IL.} Delarbre. in Stewart, Pearman & Preston 11994}, lac. cit.Chapman, S. B. 11994). Gentiana pneumonan<strong>the</strong> L. in Stewart, Pearman & Preston (19941. lac. cit.Chatters, C. (1994). Mentha pUlegium L., Moenchia erecta (L.) Gaertner, Meyer & Scherb., and Myosurus minimus L., inStewart, Pearman & Preston (1994), loc. eit.Chicken, E. 11977). Myriophyllum aquaticum IVelloso} Verdc. IM. brasi/iense Camb.} in Britain. Watsonia 11: 375-376.Clapham, A. R. (1953). Human factors contributing to a change in our flora: <strong>the</strong> former ecological status <strong>of</strong> certainhedgerow species. In Lousley, J. E., ed. The changing flora af Britain. BSSI, Oxford.Clapham, A. R., Tutin. T. G. & Warburg, E. F. 119501. <strong>Flora</strong> <strong>of</strong> <strong>the</strong> <strong>British</strong> <strong>Isles</strong>. 1st edition. Cambridge University Press,Cambridge.Clapham, A. R., Tutin. T. G. & Moore, D. M. 11987}. <strong>Flora</strong> <strong>of</strong> <strong>the</strong> <strong>British</strong> <strong>Isles</strong>. 3rd edition. Cambridge University Press,Cambridge.Clark, J. E. 11903}. Excursion Report. June 26th.- To <strong>Ashdown</strong> <strong>Forest</strong>. Proceedings <strong>of</strong> Croydon Natural History andScientific <strong>Society</strong> 5: 126-127.Clement, E. J. & Foster, M. C. (1994). Alien plants <strong>of</strong> <strong>the</strong> <strong>British</strong> <strong>Isles</strong>. BSBI, London.Coleman, W. H. (1836). <strong>Flora</strong> <strong>of</strong> East Grinstead. see Rich, T. C. G. 11994a}. W. H. Coleman's <strong>Flora</strong> <strong>of</strong> East Grinstead.Sussex <strong>Botanical</strong> Recording <strong>Society</strong>. East Grinstead.Coleman, W. H. 11849}. No. 2910. Carex boenninghausiana. In, Smith, J. E. & Sowerby, J., Supplement to EnglishBotany. 1849. London.Coles, S. M. (1977). Ranunculus rep ens L. in Europe. Watsania 11: 353-366.
236Conally, A. P. (1977). The distribution and history in <strong>the</strong> <strong>British</strong> <strong>Isles</strong> <strong>of</strong> some alien species <strong>of</strong> Po!ygonum and Reynoutria,Watsonia 11 : 291-311.Cook, C. D. K. (1966). Studies in Ranunculus subgenus Batrachium. Ill. Ranunculus hederaceus L. and R. omiophy/lus Ten.Watsonia 6: 246-259.Cook, C. D. K. & Nicholls, S. M. (1986). A monographic study <strong>of</strong> <strong>the</strong> genus Sparganium (Sparganiaceae). Part 1.Subgenu5 Xanthosparganium Holmberg. Botanica Helvetica 96: 213-267.Cook. C. D. K. (1962). Biological <strong>Flora</strong> <strong>of</strong> <strong>the</strong> <strong>British</strong> <strong>Isles</strong>. Sparganium erectum L. Journal <strong>of</strong> Ecology 82: 247-255.Cooke, A. 8., Farrell, L., Kirby, K. J. & Thomas, R. C. (1995). Changes in abundance and size <strong>of</strong> dog's mercury apparentlyassociated with grazing by muntjac. Deer 9: 429~433,Coo mbe, D. E. (1956). Biological <strong>Flora</strong> <strong>of</strong> <strong>the</strong> <strong>British</strong> <strong>Isles</strong>. Impatiens parviflora DC. Journal <strong>of</strong> Ecology 44: 701-713.Cooper, T. H. (1835). Appendix 11. Botany. In, Horsefeld, T. W. (1835). The history, antiquities and topography <strong>of</strong> <strong>the</strong>county <strong>of</strong> Sussex. Lewes.Cooper, T. H. (1862). The Botany <strong>of</strong> <strong>the</strong> County <strong>of</strong> Sussex. Baxter, Lewes.Crackles, E. F. (1990). Hypericum x desetangsii Lamotte nm. desetangsii in Yorkshire, with special reference to its spreadalong railways. Watsonia 18: 63~67.Craddock, W. A. (1953). The deer <strong>of</strong> <strong>Ashdown</strong> <strong>Forest</strong>. Sussex County Magazine 27: 294.Crowder, A. A., Pearson, M. C., Grubb, P. J. & Langlois, P. H. (1990). Biological <strong>Flora</strong> <strong>of</strong> <strong>the</strong> <strong>British</strong> <strong>Isles</strong>. Drosera L.Journal <strong>of</strong> Ecology 78: 233-267.Dale, M. P. & Causton, D. R. (1992a). The ecophysiology <strong>of</strong> Veronica chamaedrys, V. montana and V. <strong>of</strong>ficinalis. 11. Theinteraction <strong>of</strong> irradiance and water regime. Journal <strong>of</strong> Ecology 80: 493-504.Dale, M. P. & Causton, D. R. (1992b). The ecophysiology <strong>of</strong> Veronica chamaedrys, V. montana and V. <strong>of</strong>ficinalis. Ill.Effects <strong>of</strong> shading on biomass allocations - a field experiment. Journal <strong>of</strong> Ecology 80: 505-515.Darwin, C. (1867). On <strong>the</strong> specific differences between Primula veris, P. vulgaris and P. elatior; and on <strong>the</strong> hybrid nature<strong>of</strong> <strong>the</strong> common oxlip. <strong>Botanical</strong> Journal <strong>of</strong> <strong>the</strong> Unnean <strong>Society</strong> 10: 437-454.Darwin, C. (1882). The power <strong>of</strong> movement in plants. John Murray, London.Davison, A. W. (1971). The ecology <strong>of</strong> Hordeum murinum L. 11. The ruderal habitat. Journal <strong>of</strong> Ecology 59: 493-506.Davy, A. J. (1980). Biological <strong>Flora</strong> <strong>of</strong> <strong>the</strong> <strong>British</strong> <strong>Isles</strong>. Deschampsia caespitosa (L.) Beauv. Journal <strong>of</strong> Ecology 68: 1075-1096.Deakin, R. (1871). The flowering plants <strong>of</strong> Tunbridge Wells and neighbourhood. Stidolph & Bellamy, Tunbridge Wells.Dent, G. (1927-1953). Unpublished personal diary, currently held by Mrs. P. Green and to be deposited in <strong>the</strong> NaturalHistory Museum.Done, W. E. P. (1914). <strong>Flora</strong> <strong>of</strong> Groombridge, Sussex and neighbourhood recorded from personal observation in <strong>the</strong> period1904-1914. Unpublished personal diary.Dony, J. G. (1963). The expectation <strong>of</strong> plant records from prescribed areas. Watsonia 5: 377-385.Dony, J. G. (1976). Species area relationship in Bedfordshire. Bedfordshire Naturalist 30: 61-62.Drummond, J. C. & Wilbraham, A. (1939). The Englishman's food. Jonathan Cape, London.Dudman, A. A. & Richards, A. J. (1995). Dandelions <strong>of</strong> <strong>the</strong> <strong>British</strong> <strong>Isles</strong>. BSBI Handbook no. 10. (Draft March 1995version). BSSI London.Earl, D. P. (1994). Wild harvest 1. Which brambles have <strong>the</strong> tastiest berries? Plants, people and places. Newsletter No.3: 6. Liverpool Museum.Edees, E. S. & Newton, A. (1988). Brambles <strong>of</strong> <strong>the</strong> <strong>British</strong> <strong>Isles</strong>. The Ray <strong>Society</strong>, London.Ellenberg, H. (1980). Vegetation ecology <strong>of</strong> central Europe. Fourth edition. Cambridge University Press, Cambridge.Ellis, R. G. (1986). Standard tetrad nomenclature. BSBI News 43: 9.Ellis, R. G. (1994). Meconopsis cambrica (L.) Viguier. In Stewart, Pearman & Preston (1994), lac. cit.Eriksson, 0. (1988). Ramet behaviour and population growth in <strong>the</strong> clonal herb Potentilla anserina. Journal <strong>of</strong> Ecology 76:522-536.Ernst, W. H. O. (1979). Population biology <strong>of</strong> Allium ursinum in Nor<strong>the</strong>rn Germany. Journal <strong>of</strong> Ecology 67: 347-362.Evans, S. B. (1994). Viola lactea Smith. In Stewart, Pearman & Preston (1994). lac. cit.Farrell, L. (1994). Gnaphalium sylvaticum L. in Stewart, Pearman & Preston (1994), lac. cit.Feldman, S. R. & Lewis, J. P. (1990). Output and dispersal <strong>of</strong> propagules <strong>of</strong> Carduus acanthoides L. Weed Research 30:161-169.Fincham, P. (1995). Flowering plants <strong>of</strong> <strong>Ashdown</strong> <strong>Forest</strong>. A pocket guide to flowering plants which may be found on<strong>Ashdown</strong> <strong>Forest</strong>. Privately published, Fairwarp.Firmin, B. (1890). An illustrated guide to Crowborough. The Hansard Publishing Union Ltd., London.Foley, M. J. y. (1994). Orobanche rapum-genistae Thuill. In Stewart, Pearman & Preston (1994). lac. cit.Forster, T. F. (1816). <strong>Flora</strong> Tonbridgensis. J. Sprauge, Tonbridge Wells.Gibson, W. (1993a). Selective advantages to hemi-parasitic annuals genus Melampyrum <strong>of</strong> a seed dispersal mutualisminvolving ants. 1. Favourable nest sites. Oikos 67: 334-344.Gilbert, O. L. (1995). Biological <strong>Flora</strong> <strong>of</strong> <strong>the</strong> <strong>British</strong> <strong>Isles</strong>. Symphoricarpos albus (L.) S. F. Blake. Journal <strong>of</strong> Ecology 83:159-166.Gimmingham, C. H. (1960). Biological <strong>Flora</strong> <strong>of</strong> <strong>the</strong> <strong>British</strong> <strong>Isles</strong>. Calluna vulgaris L. Journal <strong>of</strong> Ecology 48: 455-483.Glyn, P. J. & Prendergast, H. (1995). <strong>Ashdown</strong> <strong>Forest</strong>. An illustrated guide. Essedon Press, Ashurstwood.Goulder, R. & Boatman, D. J. (1971). Evidence that nitrogen supply influences <strong>the</strong> distribution <strong>of</strong> a freshwatermacrophyte, Ceratophyllum demersum. Journal <strong>of</strong> Ecology 59: 783-791.
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Flora of Ashdown Forest,-,;I
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Copyright Tim Rich, Pat Donovan, Pa
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ACKNOWLEDGEMENTSWe would like to th
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THE HABITATS OF ASH DOWN FOREST3His
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Near the bottom of all the non~wood
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Conopodium majusCrataegus laevigata
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Fincham (1995) published a small ph
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13ANAL VS IS OF THE FLORA DATAIn th
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15The number of different visits to
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17Hedgerows Disturbed/waste ground
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19The frequencies with which specie
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21(r~0.434, pO.l).-0.059,•300t350
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23A second type of comparison has b
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25Similarly, environmental variable
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SPECIES ACCOUNTS27Lat'1n names foll
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29EQUISETACEAEEquisetum f/uviatile.
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One plant on a vertical, north-faci
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Viola pa/ustris in an open alder co
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35DRYOPTERIDACEAEPolystichum aculea
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37BLECHNACEAEBlechnum spicant. Hard
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39AngiospermsLAURACEAE* Laurus nobi
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41Ranunculus repens, Creeping butte
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wet mud, small streams, ditches and
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45*Meconopsis cambrica. Welsh poppy
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47Ulmusx hollandica f?U. glabra x m
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49FAGACEAEFagus sylvatica. Beech.Fo
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A special large oak at Chuck Hatch,
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Cory/us avel/ana. Hazel.Recorded in
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plants in the autumn; we have not r
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Sagina procumbens. Procumbent pearl
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59*Saponaria officinalis. Soapwort,
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61Dolygonum arenastrum. Equal-leave
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63Rumex conglomeratus. Clustered do
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65Hypericum humifusum. Trailing S1
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67DROSERACEAEDrosera rotundifolia.
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69Viola laetea. Pale dog-violet.Thi
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71* Salix alba. White willow.3 -+-t
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73Salix repens var. repens (var. er
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75*Rorippa sylvestris. Creeping yel
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77* Thlaspi arvense. Field pennycre
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79aphanus raphanistrum. Wild radish
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ica tetralix. Cross~leaved heath.As
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centre of the Forest on alluvial so
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It is predominantly a plant which o
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87SAXIFRAGACEAE[*Darmera pe/tata. I
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89insipid in flavour IHaskel1 1960)
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91ection Rubus. Subsection Hiemales
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93Rubus pr%ngatus.Nutley (45.27), 1
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95bus vestitus.Newbridge, 1969, B.
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97Rubus largificus.3 -+t--------'+_
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Potentilla anserina. Silverweed.Com
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101Geum urbanum. Wood avens.Primari
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103osa x pseudorusticana fR. arvens
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105Prunus spinosa. Blackthorn, Wint
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orbus aucuparia. Rowan, Mountain as
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109r*Crataegus persimilis. Broad-le
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111icia tetrasperma. Smooth tare.Fo
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113Trifolium rep ens. White clover"
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115Genista tinctoria. Dyer's greenw
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*Genista anglica. Petty whin, Needl
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Myriophyllum alterniflorum. Alterna
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121Epilobium montanum x obscurum.Fo
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habitats.It lives up to the name of
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125,UPHORBIACEAEVlercurialis perenn
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Although one of Britain's tiniest p
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129*Oxalis articulata. Pink-sorrel.
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*Erodium cicutarium. Common stork's
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Widespread in Europe from the shore
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* Sison amomum. Stone parsley.Tetra
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137Centaurium pu/chellum. Lesser ce
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139CONVOLVULACEAEConvolvulus arvens
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141*Nymphoides pe/tata. Fringed wat
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Myosotis secunda (M. repens). Creep
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145Stachys 8lvensis. Field woundwor
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5cutellaria x hybrida (5. galericul
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Origanum vu/gare. Marjoram.Near Fis
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CALLlTRICHACEAE151Ca/litriche is a
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153Fraxinus excelsior. Ash.4 5Local
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~'Cymbalaria mural is. Ivy-leaved t
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157Veronica scutellata. Marsh speed
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159Sibthorpia europaea. Cornish mon
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161OROBANCHACEAEOrobanche rapum~gen
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163RUBIACEAESherardia arvensis. Mad
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165Viburnum OpUIU5.Guelder-rose, St
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167Valeriana officina/is 5.1. Commo
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169Cirsium vu/gate. Spear thistle.W
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171We suspect that Leontodon hispid
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173Mycelis muralis. Wall lettuce.Se
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175Taraxacum exceJlens.Scattered al
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177Taraxacum sinuatum.Recorded from
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179Crepis capillaris. Smooth hawk'
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181Hieracium. Hawkweeds.3134 5Hiera
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*So/idago gigantea. Early goldenrod
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- Page 192 and 193: *Senecio x albescens (S. cineraria
- Page 194 and 195: *Petasites japonicus. Giant butterb
- Page 196 and 197: *E1odea canadensis. Canadian waterw
- Page 198 and 199: Arum maculatum. Lords~and~ladies, C
- Page 200 and 201: Juncus x surrejanus (J. acutiflorus
- Page 202 and 203: fences or topography. It may domina
- Page 204 and 205: Eleocharis multicaulis. Many-stalke
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- Page 208 and 209: 203Carex riparia. Greater pond~sedg
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- Page 212 and 213: 207Festuca pratensis. Meadow fescue
- Page 214 and 215: 209Cynosurus cristatus. Crested dog
- Page 216 and 217: 211Glyceria f/uitans. Floating swee
- Page 218 and 219: 213Deschampsia cespitosa. Tufted ha
- Page 220 and 221: Agrostis gigantea. Black bent.Tetra
- Page 222 and 223: 217Bromus lepidus. Slender soft~bro
- Page 224 and 225: * Triticum aestivum. Bread wheat.A
- Page 226 and 227: 221L1L1ACEAENarthecium ossifragum.
- Page 228 and 229: 223*Allium triquetrum. Three-Gornar
- Page 230 and 231: Trumpet length(mm)454035302520•
- Page 232 and 233: 227ORCHIDACEAEEpipactis purpurata.
- Page 234 and 235: een unable to refind any bog orchid
- Page 236 and 237: 231DactylorhizB praetermissa. South
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- Page 244 and 245: Rich, T. C. G. & Smith, P. A. (1996
- Page 246 and 247: GAZETTEER241This gazetteer has been
- Page 248 and 249: Smockfarthing 40.31Smuggler's car p
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- Page 252 and 253: Broad bean 111 Cardamine x hausskne
- Page 254 and 255: Fen bedstraw 163 Fumitory 46 Greate
- Page 256 and 257: Leucanthemum maximum 186 Many-flowe
- Page 258 and 259: Quercus ilex 50 Rorippa x ster/lis
- Page 260 and 261: Sticky mouse-ear 56 Thale cress 73