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Flora of Ashdown Forest - Botanical Society of the British Isles

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160Euphrasia nemorosa. Eyebright.Common (Hall 1980).This is <strong>the</strong> commonest eyebright on <strong>the</strong> <strong>Forest</strong>, widely spread along verges, rides and ingrassland. Plants with pinky~purple flowers occur in a number <strong>of</strong> populations (e.g. Millbrook4 5Hill, Millbrook Farm, Duddleswell) which have been suggested to be E. micrantha (alsoreported from Crowborough Common), but <strong>the</strong>y o<strong>the</strong>rwise conform to E. nemorosa and arebest treated as that (A. J. Silverside, pers. comm. 1994). Whe<strong>the</strong>r <strong>the</strong> flower colouration isrelict from hybridisation with E. micrantha or not remains to be seen.Widespread in England and Wales but becoming scarce in Scotland, Endemic to northand central Europe as far south as <strong>the</strong> north-east <strong>of</strong> Spain.Odontites vernus. Red bartsia.We have significantly more records than Hall (1980) who shows it in only four tetrads in <strong>the</strong><strong>Forest</strong>, probably due to our more intensive coverage <strong>of</strong> <strong>the</strong> area. It is found in a variety <strong>of</strong>habitats, in grassy areas, waste places, by tracks and rides. We have not investigated whichsubspecies occur.Widespread in Britain, and in most <strong>of</strong> Europe and nor<strong>the</strong>rn Asia.J ++-...... "------""--'-If-4 5Rhinanthus minor. Yellow~rattle, Penny~rattle.Only recorded from two places in <strong>the</strong> east <strong>of</strong> <strong>the</strong> <strong>Forest</strong>: verges by Gills Lap car park(466.315), 1987, 1993, PW; King's Standing (473.301), 1995, <strong>Flora</strong> meeting. It may be on<strong>the</strong> verges where limestone dust from <strong>the</strong> roads has spread making <strong>the</strong> soils more calcareous.It is a lot scarcer than would be expected from <strong>the</strong> distribution shown in Hall (1980) who hasJ -++-~~--"'--L+recordsfrom six tetrads, none <strong>of</strong> which correspond to our records.R. minor is indicative <strong>of</strong> undisturbed grassland and is still common on chalk in Sussex butbecoming scarcer elsewhere. The reason for <strong>the</strong> apparent decline in <strong>the</strong> last 20-30 years maybe <strong>the</strong> continued loss <strong>of</strong> old grassland to agricultural improvements.4 5This is ano<strong>the</strong>r annual hemi~parasite whose roots have been recorded attached to over50 different hosts, mostly grasses and legumes. It <strong>of</strong>ten occurs in patches and <strong>the</strong> areas around <strong>the</strong> plants have a shorter,more open sward structure and tewer grasses due to <strong>the</strong> effect it has on its hosts. It is also sensitive to grazing and canbe rapidly eliminated from swards, where it would o<strong>the</strong>rwise become a pest.It is a very variable species and is widespread in Britain and Europe. It also occurs in Greenland and North America.Pedicularis palustris. Marsh lousewort.Fisher's Bog [probably near Fisher's Gate], Or Milner Barry (Deakin 1871). <strong>Ashdown</strong> <strong>Forest</strong>, TWNH (Wolley-Dod 1937).Not seen during <strong>the</strong> survey for Hall (1980) and last seen in Sussex on <strong>the</strong> Lewes Levels, 1932, Ran Boniface (<strong>the</strong>record Willard 1989 is an error for P. sylvatica). It is a species <strong>of</strong> wet mineral or peaty flushes. Decreasing in England(Rich & Woodruff 1996), but still reasonably widespread in south-west Surrey, <strong>the</strong> New <strong>Forest</strong> and north-west Britain. It isquite widespread in Europe.Pedicularis sylvatica subsp. sylvatica. Lousewort.Common on <strong>the</strong> <strong>Forest</strong> (Hall 1980).Our distribution map shows it to be mainly concentrated on <strong>the</strong> sou<strong>the</strong>rn slopes <strong>of</strong> <strong>the</strong><strong>Forest</strong> in open areas. It is locally frequent on rides and grassy places on heathy soils, <strong>the</strong> pinkflowers a pleasant sight in early summer. All material examined has had glabrous calyces andis subsp. sylvatica. It is a biennial which is <strong>of</strong>ten abundant after fires.[t is ano<strong>the</strong>r hemi-parasite, thought to parasitise <strong>the</strong> roots <strong>of</strong> grasses. MM has carriedout experiments on its germination (Pedicularis has <strong>the</strong> reputation <strong>of</strong> being almost impossibleto grow from seed). P. sy!vatica was found to germinate readily but <strong>the</strong> numerous seedlingsdied gradually. In only one case, using Agrostis as <strong>the</strong> host, has a plant been produced whichhas flowered.Widespread and locally common in <strong>the</strong> north and west <strong>of</strong> Britain. It is decreasing in England (Rich & Woodruff 1996).Subsp. sy/vatica is endemic to Europe, and occurs in west and central Europe from Spain to Sweden and Russia.

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