*Genista anglica. Petty whin, Needle furze.On <strong>Ashdown</strong> <strong>Forest</strong> (Col em an 18361. <strong>Ashdown</strong> <strong>Forest</strong> 1904, J. Stirling (TLS). Near Gills Lap;Friar's Gate (Done 1914). Frequent on <strong>the</strong> <strong>Forest</strong> (Hall 1980).Concentrated on <strong>the</strong> east side <strong>of</strong> <strong>the</strong> <strong>Forest</strong> in <strong>the</strong> Gills Lap to Camp Hill area (e,g.abundant on rides near Stone Cottage), and also elsewhere where <strong>the</strong>re is good heath land .<strong>Ashdown</strong> <strong>Forest</strong> is <strong>the</strong> centre <strong>of</strong> its distribution in Sussex, and it is surprisingly rare on <strong>the</strong>Lower Greensand in West Sussex, Like many o<strong>the</strong>r plants <strong>of</strong> lowland heaths it is decreasing inEngland (Rich & Woodruff 1996)' but it is still scattered on upland moorland in easternScotland. It occurs in western Europe from Sweden to Italy, and in north-west Africa.Be warned, plants may sometimes be spineless (var. subinermis) and sometimes hairy ...which does not make <strong>the</strong>m G, pilosa; we have not as yet noted <strong>the</strong>se variants on <strong>the</strong> <strong>Forest</strong>.V/ex europaeus, Gorse, Furze, Hawth, Hoth. (Plate 5)"Customary tenants ought by custom to have heath, bracken and gorse to burn for <strong>the</strong>irneeds" Extent <strong>of</strong> <strong>Ashdown</strong> <strong>Forest</strong>, 1274. Ubiquitous on <strong>the</strong> <strong>Forest</strong> (Hall 1980).Gorse generally tends to occur on acid brown earths ra<strong>the</strong>r than heath podzols (Proctor1994), so it is <strong>of</strong>ten abundant along roadsides, heath margins, acidic grassland and in openscrub and is one <strong>of</strong> <strong>the</strong> commonest plants on <strong>the</strong> more open areas and roadsides on <strong>the</strong><strong>Forest</strong>. A good place to see <strong>the</strong> contrast is south <strong>of</strong> Gills Lap where it lines <strong>the</strong> road but isinfrequent in <strong>the</strong> hea<strong>the</strong>r to <strong>the</strong> east and west, though it is abundant on <strong>the</strong> steep slopesaround Gills Lap itself. Soil pHs measured at eight sites range from 2.8-5.6 with a mean <strong>of</strong>4.1 (± 0.3 s.e.). In some areas it dominates whole areas and forms dense groves; plantsopposite Churlwood car park were 2.6 metres tall.In spring <strong>the</strong> flowers scent <strong>the</strong> air with coconut and create swa<strong>the</strong>s <strong>of</strong> yellow across <strong>the</strong> <strong>Forest</strong>. Later <strong>the</strong> ripe podssplitting open with a sharp snap to scatter <strong>the</strong> seeds is a familiar sound on sunny days in July. The flowering <strong>of</strong> gorse isnotoriously variable from year to year. 1994 was a poor year for flowers, but in 1995 it started flowering pr<strong>of</strong>usely inJanuary, which was very early and continued until mid May. If it flowers early, does it set seed?At monthly intervals in spring 1995, PA and TR labelled ten flowers on each <strong>of</strong> 21 plants and followed <strong>the</strong>m throughto fruit-set. Quite a lot <strong>of</strong> <strong>the</strong> earlier flowers aborted but some did set fruit - significantly fewer in February and March thanlater. Only <strong>the</strong> February flowers had lower numbers <strong>of</strong> seeds in each pod, so even if kissing is in season, reproduction istoo!However, with <strong>the</strong> exception <strong>of</strong> those February flowers most <strong>of</strong> <strong>the</strong> seeds in <strong>the</strong> pods were predated by ei<strong>the</strong>r <strong>the</strong>gorse weevil (Apion ulicis) or a micro-moth. These seed predators are easily seen in nearly ripe pods - <strong>the</strong> weevils aregreyish and have long snouts, <strong>the</strong> micro-moth larvae are little caterpillars. Once predation has been taken into account, <strong>the</strong>actual percentage fruit set varied surprisingly little. The weevils cannot escape from <strong>the</strong> pods <strong>the</strong>mselves, and have towait until <strong>the</strong>y split open - so <strong>the</strong> familiar crack <strong>of</strong> pods may be more usually indicative <strong>of</strong> weevils flying through <strong>the</strong> airthan seeds.Summary <strong>of</strong> fruit and seed set in V/ex europaeus 'In 1995 near Long car park (10 flowers were marked on each <strong>of</strong> 21bushes at approximately four week intervals).117Date flowers markedNumber <strong>of</strong> bushes floweringMean number <strong>of</strong> pods harvested per bushMean number <strong>of</strong> seeds per un predated podMean percentage pods predatedMean number <strong>of</strong> good seeds per podMean chance <strong>of</strong> ovule producing seed4 Feb.63.21.85.3%1.85.5%5 March123.34.576%2.24.4%2 April 29 April20 216.2 5.34.5 4.661% 64.5%2.3 1.910.8% 8,1%Gorse was widely used on <strong>the</strong> <strong>Forest</strong> for ro<strong>of</strong>ing, fuel, cooking, bread making, and brick and lime kilns (Steele 1936;Leppard 1980; Irons 1982). The spines protect it to some extent from grazing, and <strong>the</strong> plant was crushed in a special millso it could be fed to animals. O<strong>the</strong>rs note that <strong>the</strong> thorns are remarkably persistent in <strong>the</strong> skin, some taking six days to getout. Blooms are still collected on <strong>the</strong> <strong>Forest</strong> to make wine, and <strong>the</strong>re were authorised sales by <strong>the</strong> Wood Reeve <strong>of</strong> gorsefor horse jumps and for planting. Arthur (1989) records that sticks <strong>of</strong> gorse from <strong>Ashdown</strong> <strong>Forest</strong> were sold for umbrellahandles. A very readable account <strong>of</strong> gorse, including <strong>the</strong> economic uses and folklore, is given by Humphries &Shaughnessy (1987).Gorse is a predominantly western plant in Europe, but is widespread in Britain. It is also widespread in Australasia asan introduction and is a noxious weed. The potential for controlling it using <strong>the</strong> weevils has been investigated but has metwith little success.
118U/ex gal/ii. Western gorse.Kidd's Hill, A. H. Wolley-Dod and W. Ingwersen IWolley-Dod 19371. Introduced to <strong>the</strong> Wych Cross area in <strong>the</strong> 1960s byM. Woodman as an experiment, but not currently known.This species occurs widely in <strong>the</strong> north and west <strong>of</strong> Britain and just reaches West Sussex, but also occurs in Kent andEast Anglia, Endemic to western Europe form Spain to Scotland. It is somewhat sensitive to frost.Care needs to be taken in separating it from U. minor; U. gallH populations usually have a mean calyx length <strong>of</strong> morethan 10 mm, whilst U. minor populations have <strong>the</strong>m less than 9 mm.V/ex minor (V. nanus). Dwarf gorse, Dog furze.<strong>Ashdown</strong> <strong>Forest</strong> IColeman 18361. On <strong>the</strong> <strong>Forest</strong>s around Crowborough IDone 19141.Abounds on some <strong>of</strong> <strong>the</strong> heaths in East Sussex, such as <strong>Ashdown</strong> <strong>Forest</strong> (Wolley-Dod 1937),Common on <strong>the</strong> <strong>Forest</strong> IHall 1980).On <strong>the</strong> <strong>Forest</strong>, it occurs mainly on dry heath with Cal/una, and on wet podzolisedheathland soils with Molinia and Erica tetra/ix. The soil pHs <strong>of</strong> seven sites measured rangefrom 3.2-4.1, with a mean <strong>of</strong> 3.7 I± 0.13 s.e.l. It grows predominantly in <strong>the</strong> open, but willsurvive for a few years in open secondary woodland where it can grow to 1.5 metres tall.Our map shows it from all squares with significant areas <strong>of</strong> heathland, and it is only absentfrom some areas around <strong>the</strong> edge. Elsewhere in Britain it occurs on chalk 'heaths' and sandyglacial drift (Proctor 1994). It is palatable and is suppressed by heavy grazing.It flowers pr<strong>of</strong>usely in August and September, <strong>the</strong> deep yellow flowers contrasting beautifully with <strong>the</strong> purple hea<strong>the</strong>r.The fruits do not appear to be predated by weevils, and <strong>the</strong>y over-winter and ripen in <strong>the</strong> spring (this seems to be veryunusual in Br'itish plants). Seeds are slow and erratic to germinate, but seedling establishment can take place rapidly afterfire.This plant is one <strong>of</strong> <strong>the</strong> real treasures <strong>of</strong> lowland heaths in <strong>the</strong> south-east <strong>of</strong> England, with a few scattered localities in<strong>the</strong> East Midlands, North Wales, Lancashire and Cumbria (Proctor 1994). The populations on <strong>the</strong>se heaths represent amajor part <strong>of</strong> <strong>the</strong> world population as, outside England, it is confined to <strong>the</strong> western edge <strong>of</strong> France, Spain and Portugal,but gives way to U. gallii along <strong>the</strong> Atlantic margin.The <strong>British</strong> distribution <strong>of</strong> U. minor scarcely overlaps at all with that <strong>of</strong> its close relative U. ga//iJ~ western gorse, whichhas a similar ecological niche.HALORAGACEAE[Myriophyl/um verticil/atum. Whorled water-milfoi!. Pond on <strong>Ashdown</strong> <strong>Forest</strong>, E. Jenner IArnold 18871. A surprisingrecord as it tends to occur in base-rich water and probably an error? The whereabouts <strong>of</strong> Jenner's herbarium is unknownso <strong>the</strong> record cannot be checked.]*Myriophyllum aquaticum (M, brasiliense), Parrot·s-fea<strong>the</strong>r.Tetrad 42U IHall 19801. Duddleswell, 1976, C. T. Prime, 1978 J. Milner IMilner 19791.Planted around pond at Chelwood Vachery 1431.297), 1994, TR; pond on golf course,<strong>Forest</strong> Row 1436.342), 1995, TR, and one <strong>of</strong> <strong>the</strong> few plants to survive bulldozing inSeptember 1995; pond by Prickets Hatch 1443.2701, TR & PA, 1995; Ellison's Pond,3 -++_~----L+_ Duddleswell 1462.2871, 1986 +, DB & B. Wurzell, growing to practically fill <strong>the</strong> pond by 19954 5- this may be one <strong>of</strong> <strong>the</strong> longest established sites in Britain.This invasive plant, which first became naturalized in Britain sometime in <strong>the</strong> early-1970s(Chicken 1977) as a throw-out from aquarists' ponds or with tadpoles or terrapins, is nowbecoming widely established in Britain. It may even be able to out-compete Crassula helmsiiby over-growing it. It is reputed to be frost-sensitive but has survived freezing conditions on <strong>Ashdown</strong> <strong>Forest</strong> (Milner1979, Hall 1980) and has certainly thrived in <strong>the</strong> run <strong>of</strong> mild winters. PW has also cultivated material from Ellison's Pondand observed that it is tolerant <strong>of</strong> cold; shoots were blackened by frost but in <strong>the</strong> spring put up new green shoots. Wesuggest it should be eradicated as soon as possible after it has been discovered.It is native in South America.Myriophyllum spicatum, Spiked water-milfoil.Pond on <strong>Ashdown</strong> <strong>Forest</strong> near Wych Cross IWhitwell 19021.Both sides <strong>of</strong> ornamental pond at <strong>Ashdown</strong> Park Hotel 1431.320), 1995, TR & PA, onesite but two squares, and interestingly in <strong>the</strong> same areas as Whitwell's record,In Sussex it occurs predominantly near <strong>the</strong> coast in streams, lakes, ponds and ditches,3 -++_------!..+_ sometimes in quantity (Hall 1980). It is widespread in Britain and is possibly commonest in<strong>the</strong> south and east, where it occurs in base-rich, moderately nutrient-rich water. Widespreadin Europe, Asia, North Africa and North America.4 5
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Flora of Ashdown Forest,-,;I
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Copyright Tim Rich, Pat Donovan, Pa
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ACKNOWLEDGEMENTSWe would like to th
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THE HABITATS OF ASH DOWN FOREST3His
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Near the bottom of all the non~wood
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Conopodium majusCrataegus laevigata
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Fincham (1995) published a small ph
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13ANAL VS IS OF THE FLORA DATAIn th
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15The number of different visits to
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17Hedgerows Disturbed/waste ground
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19The frequencies with which specie
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21(r~0.434, pO.l).-0.059,•300t350
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23A second type of comparison has b
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25Similarly, environmental variable
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SPECIES ACCOUNTS27Lat'1n names foll
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29EQUISETACEAEEquisetum f/uviatile.
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One plant on a vertical, north-faci
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Viola pa/ustris in an open alder co
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35DRYOPTERIDACEAEPolystichum aculea
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37BLECHNACEAEBlechnum spicant. Hard
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39AngiospermsLAURACEAE* Laurus nobi
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41Ranunculus repens, Creeping butte
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wet mud, small streams, ditches and
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45*Meconopsis cambrica. Welsh poppy
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47Ulmusx hollandica f?U. glabra x m
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49FAGACEAEFagus sylvatica. Beech.Fo
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A special large oak at Chuck Hatch,
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Cory/us avel/ana. Hazel.Recorded in
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plants in the autumn; we have not r
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Sagina procumbens. Procumbent pearl
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59*Saponaria officinalis. Soapwort,
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61Dolygonum arenastrum. Equal-leave
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63Rumex conglomeratus. Clustered do
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65Hypericum humifusum. Trailing S1
- Page 72 and 73: 67DROSERACEAEDrosera rotundifolia.
- Page 74 and 75: 69Viola laetea. Pale dog-violet.Thi
- Page 76 and 77: 71* Salix alba. White willow.3 -+-t
- Page 78 and 79: 73Salix repens var. repens (var. er
- Page 80 and 81: 75*Rorippa sylvestris. Creeping yel
- Page 82 and 83: 77* Thlaspi arvense. Field pennycre
- Page 84 and 85: 79aphanus raphanistrum. Wild radish
- Page 86 and 87: ica tetralix. Cross~leaved heath.As
- Page 88 and 89: centre of the Forest on alluvial so
- Page 90 and 91: It is predominantly a plant which o
- Page 92 and 93: 87SAXIFRAGACEAE[*Darmera pe/tata. I
- Page 94 and 95: 89insipid in flavour IHaskel1 1960)
- Page 96 and 97: 91ection Rubus. Subsection Hiemales
- Page 98 and 99: 93Rubus pr%ngatus.Nutley (45.27), 1
- Page 100 and 101: 95bus vestitus.Newbridge, 1969, B.
- Page 102 and 103: 97Rubus largificus.3 -+t--------'+_
- Page 104 and 105: Potentilla anserina. Silverweed.Com
- Page 106 and 107: 101Geum urbanum. Wood avens.Primari
- Page 108 and 109: 103osa x pseudorusticana fR. arvens
- Page 110 and 111: 105Prunus spinosa. Blackthorn, Wint
- Page 112 and 113: orbus aucuparia. Rowan, Mountain as
- Page 114 and 115: 109r*Crataegus persimilis. Broad-le
- Page 116 and 117: 111icia tetrasperma. Smooth tare.Fo
- Page 118 and 119: 113Trifolium rep ens. White clover"
- Page 120 and 121: 115Genista tinctoria. Dyer's greenw
- Page 124 and 125: Myriophyllum alterniflorum. Alterna
- Page 126 and 127: 121Epilobium montanum x obscurum.Fo
- Page 128 and 129: habitats.It lives up to the name of
- Page 130 and 131: 125,UPHORBIACEAEVlercurialis perenn
- Page 132 and 133: Although one of Britain's tiniest p
- Page 134 and 135: 129*Oxalis articulata. Pink-sorrel.
- Page 136 and 137: *Erodium cicutarium. Common stork's
- Page 138 and 139: Widespread in Europe from the shore
- Page 140 and 141: * Sison amomum. Stone parsley.Tetra
- Page 142 and 143: 137Centaurium pu/chellum. Lesser ce
- Page 144 and 145: 139CONVOLVULACEAEConvolvulus arvens
- Page 146 and 147: 141*Nymphoides pe/tata. Fringed wat
- Page 148 and 149: Myosotis secunda (M. repens). Creep
- Page 150 and 151: 145Stachys 8lvensis. Field woundwor
- Page 152 and 153: 5cutellaria x hybrida (5. galericul
- Page 154 and 155: Origanum vu/gare. Marjoram.Near Fis
- Page 156 and 157: CALLlTRICHACEAE151Ca/litriche is a
- Page 158 and 159: 153Fraxinus excelsior. Ash.4 5Local
- Page 160 and 161: ~'Cymbalaria mural is. Ivy-leaved t
- Page 162 and 163: 157Veronica scutellata. Marsh speed
- Page 164 and 165: 159Sibthorpia europaea. Cornish mon
- Page 166 and 167: 161OROBANCHACEAEOrobanche rapum~gen
- Page 168 and 169: 163RUBIACEAESherardia arvensis. Mad
- Page 170 and 171: 165Viburnum OpUIU5.Guelder-rose, St
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167Valeriana officina/is 5.1. Commo
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169Cirsium vu/gate. Spear thistle.W
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171We suspect that Leontodon hispid
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173Mycelis muralis. Wall lettuce.Se
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175Taraxacum exceJlens.Scattered al
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177Taraxacum sinuatum.Recorded from
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179Crepis capillaris. Smooth hawk'
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181Hieracium. Hawkweeds.3134 5Hiera
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*So/idago gigantea. Early goldenrod
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ecorded with Chamaemelum are Matric
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*Senecio x albescens (S. cineraria
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*Petasites japonicus. Giant butterb
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*E1odea canadensis. Canadian waterw
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Arum maculatum. Lords~and~ladies, C
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Juncus x surrejanus (J. acutiflorus
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fences or topography. It may domina
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Eleocharis multicaulis. Many-stalke
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A hybrid first described by Col em
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203Carex riparia. Greater pond~sedg
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Material which initially looked lik
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207Festuca pratensis. Meadow fescue
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209Cynosurus cristatus. Crested dog
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211Glyceria f/uitans. Floating swee
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213Deschampsia cespitosa. Tufted ha
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Agrostis gigantea. Black bent.Tetra
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217Bromus lepidus. Slender soft~bro
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* Triticum aestivum. Bread wheat.A
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221L1L1ACEAENarthecium ossifragum.
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223*Allium triquetrum. Three-Gornar
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Trumpet length(mm)454035302520•
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227ORCHIDACEAEEpipactis purpurata.
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een unable to refind any bog orchid
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231DactylorhizB praetermissa. South
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Rachel A. Nicholson (1934-). Rachel
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235REFERENCESAbbas, M. H. (1979). S
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Graham, G. G. & Primavesi, A. L. (1
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Rich, T. C. G. & Smith, P. A. (1996
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GAZETTEER241This gazetteer has been
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Smockfarthing 40.31Smuggler's car p
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Asple adi+cuttr int~~~tl·Bteel SI!
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Broad bean 111 Cardamine x hausskne
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Fen bedstraw 163 Fumitory 46 Greate
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Leucanthemum maximum 186 Many-flowe
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Quercus ilex 50 Rorippa x ster/lis
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Sticky mouse-ear 56 Thale cress 73