93Rubus pr%ngatus.Nutley (45.27), 1995, RR & PH, known in <strong>the</strong> area (Edees & Newton 1988) and seen atChailey Common in 1994, AN and BSBI, and Tunbridge Wells, 1995, RR.Woods, heaths and hedge banks, widespread in south-western Britain and also found innorth-western France,Rubus subinermoides (R. pubescens var. subinermis).One <strong>of</strong> <strong>the</strong> commonest Rubi on <strong>Ashdown</strong> <strong>Forest</strong>, H. J. Riddelsdell; near Wych Cross, T. Hilton(Wolley-Dod 1937). Newbridge, 1969, B. A. Miles (Philp 1970). TQ/4.3 (Hall 1980; Edees &Newton 1988).J -++-------'-t-Woods near Priory Road 142.33), Goat car park (403.326), Warren car park (416.320),opposite Wych Cross Nurseries (421.321), <strong>Ashdown</strong> <strong>Forest</strong> Visitor Centre (43.32),4 5Townsend's car park (445.328), Newbridge (456.3251. and Pooh car park to Pooh Bridge(46.33,47.33), 1994, DE. Near Coldharbour (40.32), Hindleap Warren (41.32), Coleman'sHatch (44.321. Gills Lap (46.31), roadsides near Sunting's, Newbridge (46.321. 1994, AN &AB. Nutley 144.27, 45.27), Fairwarp 146.26, 47.26), near Ellison's Pond (46.28), 1995, RR& PH.Often abundant in Sussex, and common and widespread in sou<strong>the</strong>rn England, yet endemic to <strong>the</strong> <strong>British</strong> <strong>Isles</strong>,Series Sprenge/iani.Rubus brevistaminosus.Roadsides near Sunting's, Newbridge (46.32), 1994, AN & AB (Edees & Newton 1988).A local endemic species apparently confined to <strong>the</strong> Weald and North Downs. There areseveral colonies in <strong>the</strong> Hartfield area.Rubus sprenge/ii.J -++-------'-t-TQ/4.3 (Hall 1980; Edees & Newton 1988).Warren car park (416.320), Piglet's car park (470.320), Po oh car park (472.332),1994,DE. Hindleap Warren (41.32), Stonehill (45.28), 1994, AN & AB. Nutley (44.27, 45.27,45.28), Fairwarp (46.26), 1995, RR & PH.This dainty pink-flowered bramble may be quite widespread in <strong>the</strong> <strong>Forest</strong>, Widespread incentral and nor<strong>the</strong>rn Europe, especially in oak-birch woods on sandy and gravelly soils.4 5Series Disc%res.Rubus armipotens (R. godronii, R. pseudobifrons).Near Wych Cross, W. C. Barton & H. J. Riddelsdell (Wolley-Dod 1937). Newbridge, 1969, B.A. Miles IPhilp 1970). TQ/4.3 (Hall 1980; Edees & Newton 1988).Priory Road near Kidbrooke Wood (412.3381. Goat car park (403.326), and path to Pooh3 -++---------'_+_Bridge (472.332), 1994, DE. Near Cold harbour (40.32), Hindleap Warren (41.32), Gills Lap(46.31 I. 1994, AN & AB. Nutley (44.27, 45.27, 45.28), near Ellison's Pond 146.28),4 5Fairwarp Farm (47.26), near Poundgate car park (481.284), 1995, RR & PH.An endemic species <strong>of</strong> heaths, wood margins and hedge banks, and particularly commonin south-east England.
94Rubus procerus (R. armeniacusJ. Himalayan Giant.Hindleap Warren (41.321, Stonehill (45.281, Gills Lap (46.31), 1994, AN & AB. Nutley(44.27), 1995, RR & PH.This is <strong>the</strong> cultivated blackberry which is increasingly escaping from cultivation in Britain.Despite <strong>the</strong> English name it probably originated from Germany. The berries may vary fromsweet to sour on <strong>the</strong> same panicle, though with a few ounces <strong>of</strong> sugar <strong>the</strong>y are ideal for jam'or pies (Earl 1994).Rubus ulmifolius fR. fusticanus).Newbridge, 1969, B. A. Miles (Philp 1970). T0/4.3 (Hall 1980; Edees & Newton 1988).Path to Pooh Sticks Bridge (472.332), near Townsend car park (445.328), 1994, DE.Roadsides near Sunting's, Newbridge (46.32), AN & AB, 1994. Nutley (44.27), FairwarpFarm (47.26), 1995, RR & PH.3 Perhaps one <strong>of</strong> <strong>the</strong> commonest and most widespread brambles in Br'ltain, from sea cliffsin <strong>the</strong> Channel Islands to hedges in central England, and north to Scotland, though becomingmore restricted to south~facing banks near <strong>the</strong> coast in <strong>the</strong> north.4 5 It has also been introduced throughout <strong>the</strong> world (not for its fruits which are small andsour) and is <strong>of</strong>ten a pest; for instance, it has spread dramatically on Robinson Crusoe's Islands<strong>of</strong>f <strong>the</strong> coast <strong>of</strong> Chile and is ousting <strong>the</strong> native flora which contains 41 % endemics (Plant Talk 2: 19-21).Series Vestiti.Rubus adscitus.East <strong>of</strong> Fairwarp Farm on path edge (473.266), 1995, RR & PH, <strong>the</strong> first record for EastSussex. It has long pointed sepals and floriferous zigzag panicles which make it looksomewhat more distinctive than many brambles.It is widespread in sou<strong>the</strong>rn Britain, and also occurs in France.Rubus leucostachys.3-+~----------L+-4 5Newbridge, 1969, B. A. Miles (Philp 1970). T0/4.3 (Hall 1980; Edees & Newton 1988).Woods near Priory Road (412.3381. and Townsend's car park (445.3281. 1994, DE. NearColdharbour (40.321. Hindleap Warren (41.32), Coleman's Hatch (44.32), Stonehill (45.28),Gills Lap (46.31 I. roadsides near Sunting's, Newbridge (46.32), 1994, AN & AB. Toll Lane(461.2631. Fairwarp (46.26, 47.26), near Poundgate car park (481.284),1995, RR & PH.'Var. angustata' was also recorded at Coleman's Hatch (44.32) and Stonehill (45.28),1994, AN & AB.A regional endemic species in south-east England.Rubus surrejanus.Probably <strong>Ashdown</strong> <strong>Forest</strong>, C. H. Waddell? (Wolley-Dod 1937). T0/4.3 (Hall 1980; Edees &Newton 1988). The record 'or R. lasiostachvs from Newbridge, 1969, B. A. Miles (Philp1970) refers to this species.Specimens collected at Warren car park (416.320), 1994, DE, may be this species. GillsLap (46.31), 1994, AN & AB. South <strong>of</strong> Poundgate car park (482.284), 1995, RR & PH.Hedgebanks, borders <strong>of</strong> woods and heaths, widespread and endemic in sou<strong>the</strong>rnEngland.
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Flora of Ashdown Forest,-,;I
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Copyright Tim Rich, Pat Donovan, Pa
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ACKNOWLEDGEMENTSWe would like to th
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THE HABITATS OF ASH DOWN FOREST3His
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Near the bottom of all the non~wood
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Conopodium majusCrataegus laevigata
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Fincham (1995) published a small ph
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13ANAL VS IS OF THE FLORA DATAIn th
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15The number of different visits to
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17Hedgerows Disturbed/waste ground
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19The frequencies with which specie
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21(r~0.434, pO.l).-0.059,•300t350
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23A second type of comparison has b
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25Similarly, environmental variable
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SPECIES ACCOUNTS27Lat'1n names foll
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29EQUISETACEAEEquisetum f/uviatile.
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One plant on a vertical, north-faci
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Viola pa/ustris in an open alder co
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35DRYOPTERIDACEAEPolystichum aculea
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37BLECHNACEAEBlechnum spicant. Hard
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39AngiospermsLAURACEAE* Laurus nobi
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41Ranunculus repens, Creeping butte
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- Page 50 and 51: 45*Meconopsis cambrica. Welsh poppy
- Page 52 and 53: 47Ulmusx hollandica f?U. glabra x m
- Page 54 and 55: 49FAGACEAEFagus sylvatica. Beech.Fo
- Page 56 and 57: A special large oak at Chuck Hatch,
- Page 58 and 59: Cory/us avel/ana. Hazel.Recorded in
- Page 60 and 61: plants in the autumn; we have not r
- Page 62 and 63: Sagina procumbens. Procumbent pearl
- Page 64 and 65: 59*Saponaria officinalis. Soapwort,
- Page 66 and 67: 61Dolygonum arenastrum. Equal-leave
- Page 68 and 69: 63Rumex conglomeratus. Clustered do
- Page 70 and 71: 65Hypericum humifusum. Trailing S1
- Page 72 and 73: 67DROSERACEAEDrosera rotundifolia.
- Page 74 and 75: 69Viola laetea. Pale dog-violet.Thi
- Page 76 and 77: 71* Salix alba. White willow.3 -+-t
- Page 78 and 79: 73Salix repens var. repens (var. er
- Page 80 and 81: 75*Rorippa sylvestris. Creeping yel
- Page 82 and 83: 77* Thlaspi arvense. Field pennycre
- Page 84 and 85: 79aphanus raphanistrum. Wild radish
- Page 86 and 87: ica tetralix. Cross~leaved heath.As
- Page 88 and 89: centre of the Forest on alluvial so
- Page 90 and 91: It is predominantly a plant which o
- Page 92 and 93: 87SAXIFRAGACEAE[*Darmera pe/tata. I
- Page 94 and 95: 89insipid in flavour IHaskel1 1960)
- Page 96 and 97: 91ection Rubus. Subsection Hiemales
- Page 100 and 101: 95bus vestitus.Newbridge, 1969, B.
- Page 102 and 103: 97Rubus largificus.3 -+t--------'+_
- Page 104 and 105: Potentilla anserina. Silverweed.Com
- Page 106 and 107: 101Geum urbanum. Wood avens.Primari
- Page 108 and 109: 103osa x pseudorusticana fR. arvens
- Page 110 and 111: 105Prunus spinosa. Blackthorn, Wint
- Page 112 and 113: orbus aucuparia. Rowan, Mountain as
- Page 114 and 115: 109r*Crataegus persimilis. Broad-le
- Page 116 and 117: 111icia tetrasperma. Smooth tare.Fo
- Page 118 and 119: 113Trifolium rep ens. White clover"
- Page 120 and 121: 115Genista tinctoria. Dyer's greenw
- Page 122 and 123: *Genista anglica. Petty whin, Needl
- Page 124 and 125: Myriophyllum alterniflorum. Alterna
- Page 126 and 127: 121Epilobium montanum x obscurum.Fo
- Page 128 and 129: habitats.It lives up to the name of
- Page 130 and 131: 125,UPHORBIACEAEVlercurialis perenn
- Page 132 and 133: Although one of Britain's tiniest p
- Page 134 and 135: 129*Oxalis articulata. Pink-sorrel.
- Page 136 and 137: *Erodium cicutarium. Common stork's
- Page 138 and 139: Widespread in Europe from the shore
- Page 140 and 141: * Sison amomum. Stone parsley.Tetra
- Page 142 and 143: 137Centaurium pu/chellum. Lesser ce
- Page 144 and 145: 139CONVOLVULACEAEConvolvulus arvens
- Page 146 and 147: 141*Nymphoides pe/tata. Fringed wat
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Myosotis secunda (M. repens). Creep
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145Stachys 8lvensis. Field woundwor
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5cutellaria x hybrida (5. galericul
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Origanum vu/gare. Marjoram.Near Fis
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CALLlTRICHACEAE151Ca/litriche is a
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153Fraxinus excelsior. Ash.4 5Local
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~'Cymbalaria mural is. Ivy-leaved t
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157Veronica scutellata. Marsh speed
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159Sibthorpia europaea. Cornish mon
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161OROBANCHACEAEOrobanche rapum~gen
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163RUBIACEAESherardia arvensis. Mad
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165Viburnum OpUIU5.Guelder-rose, St
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167Valeriana officina/is 5.1. Commo
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169Cirsium vu/gate. Spear thistle.W
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171We suspect that Leontodon hispid
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173Mycelis muralis. Wall lettuce.Se
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175Taraxacum exceJlens.Scattered al
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177Taraxacum sinuatum.Recorded from
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179Crepis capillaris. Smooth hawk'
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181Hieracium. Hawkweeds.3134 5Hiera
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*So/idago gigantea. Early goldenrod
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ecorded with Chamaemelum are Matric
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*Senecio x albescens (S. cineraria
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*Petasites japonicus. Giant butterb
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*E1odea canadensis. Canadian waterw
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Arum maculatum. Lords~and~ladies, C
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Juncus x surrejanus (J. acutiflorus
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fences or topography. It may domina
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Eleocharis multicaulis. Many-stalke
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A hybrid first described by Col em
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203Carex riparia. Greater pond~sedg
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Material which initially looked lik
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207Festuca pratensis. Meadow fescue
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209Cynosurus cristatus. Crested dog
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211Glyceria f/uitans. Floating swee
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213Deschampsia cespitosa. Tufted ha
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Agrostis gigantea. Black bent.Tetra
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217Bromus lepidus. Slender soft~bro
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* Triticum aestivum. Bread wheat.A
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221L1L1ACEAENarthecium ossifragum.
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223*Allium triquetrum. Three-Gornar
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Trumpet length(mm)454035302520•
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227ORCHIDACEAEEpipactis purpurata.
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een unable to refind any bog orchid
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231DactylorhizB praetermissa. South
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Rachel A. Nicholson (1934-). Rachel
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235REFERENCESAbbas, M. H. (1979). S
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Graham, G. G. & Primavesi, A. L. (1
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Rich, T. C. G. & Smith, P. A. (1996
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GAZETTEER241This gazetteer has been
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Smockfarthing 40.31Smuggler's car p
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Asple adi+cuttr int~~~tl·Bteel SI!
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Broad bean 111 Cardamine x hausskne
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Fen bedstraw 163 Fumitory 46 Greate
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Leucanthemum maximum 186 Many-flowe
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Quercus ilex 50 Rorippa x ster/lis
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Sticky mouse-ear 56 Thale cress 73