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Flora of Ashdown Forest - Botanical Society of the British Isles

Flora of Ashdown Forest - Botanical Society of the British Isles

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125,UPHORBIACEAEVlercurialis perennis. Oog's mercury, Snake's-bit, Snake's victuals.Frequent around <strong>the</strong> <strong>Forest</strong>, but notably one <strong>of</strong> <strong>the</strong> few places in Sussex where it is notubiquitous outside <strong>the</strong> urban areas (Hall 1980).Most <strong>of</strong> our records are from <strong>the</strong> more calcareous soils around <strong>the</strong> edge <strong>of</strong> <strong>the</strong> <strong>Forest</strong>and in <strong>the</strong> older woodlands, but in some places it is established in secondary habitats as on3 <strong>the</strong> A22 road embankment over Millbrook (442.286).4 5This is a rhizomatous perennial <strong>of</strong> shaded places in <strong>the</strong> lowlands on moderately tostrongly calcareous, generally freely-draining soils. Brewls, Bowman & Rose (1996) note thatit does not withstand trampling. It is one <strong>of</strong> <strong>the</strong> first plants to flower in <strong>the</strong> spring and isdioecious. The two sexes are reported to have different ecological requirements, <strong>the</strong> malesbeing more commonly recorded in well-lit places and higher pH soils (Grime et al. 1994). Five colonies investigated in <strong>the</strong>more open parts <strong>of</strong> <strong>the</strong> <strong>Forest</strong> in spring 1996 were exclusively male, suggesting colonisation from <strong>the</strong> surrounding woods.Widespread in Britain except in <strong>the</strong> far north and in Europe south <strong>of</strong> <strong>the</strong> Baltic, Caucasus and south-west Asia.*Mercurialis annua. Annual mercury.Tetrad 43V (Hall 1980). Not refound. A weed <strong>of</strong> gardens and cultivated ground usually on calcareous or nutrient-rich soilsin sou<strong>the</strong>rn Britain, but not established in <strong>the</strong> <strong>Forest</strong>. Native in Eurasia and North Africa. It is an unusual species withdioecious diploids and monoecious polypi aids; to date only <strong>the</strong> former have been recorded in Britain.Euphorbia helioscopia. Sun spurge.Tetrads 42E, 42T and 43H (Hall 1980).On <strong>the</strong> <strong>Forest</strong> it is an annual weed <strong>of</strong> arable fields and gardens. It is in five scatteredsquares and is especially associated with <strong>the</strong> villages.In Sussex widespread on <strong>the</strong> chalk in arable land, and similarly widespread in lowland3 -++_--'~----L+_Britain, Europe and central Asia.4 5*Euphorbia lathyris. Caper spurge.,-,--;:==,---,.---, Recorded once as an escape from cultivation outside houses at Streeter's Rough (423.292)'1993, TR. It was once planted on <strong>the</strong> green in front <strong>of</strong> Duddleswell tea rooms as a moledeterrent; <strong>the</strong> plants have gone but <strong>the</strong> moles obviously still like <strong>the</strong> local food source.A garden weed scattered in Sussex and increasing in England (Rich & Woodruff 1996).3 -+h..------L+_ Probably native in east and central Mediterranean region.4 5Euphorbia pep/us. Petty spurge.Common around <strong>the</strong> <strong>Forest</strong> (Hall 1980).We have also found it common around <strong>the</strong> <strong>Forest</strong> edges, <strong>the</strong> exceptions being two siteson soil introduced to Pippingford Park. It is much more frequent as an arable weed in <strong>the</strong>villages than E. helioscopia, possibly because it has broader soil requirements, which may also3 explain its wider distribution in Britain.Widespread in lowland Britain and Europe to Siberia.4 5Euphorbia amygdaloides.3 -+--ia _____ --4 5Wood spurge.Frequent around <strong>the</strong> <strong>Forest</strong> (Hall 1980).Our records largely match· <strong>the</strong> pattern in Hall (1980) where it occurs around <strong>the</strong> <strong>Forest</strong>edges with a few records in <strong>the</strong> middle. It is most frequent in <strong>the</strong> Toll Lane woodlands where<strong>the</strong> soils are more calcareous clays, but is also surprisingly frequent in parts <strong>of</strong> Pipping fordPark. It seems to be largely uneaten by deer.This is a classic plant <strong>of</strong> coppiced woodlands on moderately to strongly calcareous soHs,and is quite widespread in Sussex. It is locally frequent in sou<strong>the</strong>rn Britain north to <strong>the</strong>Midlands, and in central, sou<strong>the</strong>rn and north-western Europe, <strong>the</strong> Caucasus and <strong>the</strong> Algerianmountains.

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