Sagina procumbens. Procumbent pearlwort, Bird's-eye.Common on <strong>the</strong> <strong>Forest</strong> IHall 1980}.Recorded in all but two squares on <strong>the</strong> <strong>Forest</strong>, where it is typical <strong>of</strong> short, damp, opengrassland, lawns, car parks, and disturbed ground.Common and widespread in Sussex, Britain, Europe, Asia, North America and Greenland.3574 5Sagina apetala. Annual pearlwort.Common on <strong>the</strong> <strong>Forest</strong> IHall 1980}.We have it mainly on <strong>the</strong> west side <strong>of</strong> <strong>the</strong> <strong>Forest</strong> where it occurs on disturbed soils in carparks, where it may have been introduced with gravel, on tracks, and around <strong>the</strong> villages. Wehave recorded both subsp. apetala and subsp. erecta and agree with Hall (1980) that <strong>the</strong>3 latter is commoner, but have not looked at <strong>the</strong>ir distributions in detail.Widespread in Britain but mainly south-western in Europe, and also in western Asia.4 5Scleranthus annuus. Annual knawel.<strong>Forest</strong> Row, Miss P. Stockdale, 1910 IBEX; WolleY-Dod 1937}.Not refound on <strong>the</strong> forest, and probably declining in Sussex. It is most persistent on dry, open, light, sandy soils andmay not have liked our damp, silty soils.Scattered patchily through most <strong>of</strong> Britain on suitable acidic soils but decreasing (Rich & Woodruff 1996), thoughwidespread in Europe, Asia and North Africa.Spergula arvensis. Corn spurrey.Nine tetrads in Hall 11980}.We only have five records from open disturbed ground, and it does seem rare.It is a phenotypically and genetically variable species. In Britain <strong>the</strong> two main varietiesshow an interesting difference in distribution, which New & Herriott (1981) ascribe to3 -++-_.!IL.._--'"JLjdifferentgermination characteristics; plants examined from two sites on <strong>the</strong> <strong>Forest</strong> were var.arvensis which lacks papillae and is <strong>the</strong> commoner one in south-east England, whilst var.sativa which has papillae occurs more frequently in <strong>the</strong> north and west.Locally frequent on sandy soils in lowland Britain but decreasing (Rich & Woodruff 1996)4 5and widespread in Europe except around <strong>the</strong> Mediterranean. It is cosmopolitan indistribution and reportedly one <strong>of</strong> <strong>the</strong> world's worst weeds, especially in cereal crops.*Spergularia marina. Lesser sea-spurrey.One plant <strong>of</strong> this seaside plant was found on bare soil on <strong>the</strong> A22 verge at Kidbrooke Hill1418.335}, 1995, TR & PA.As a native it occurs on salt marshes and saline places around <strong>the</strong> coast <strong>of</strong> Britain andEurope and <strong>the</strong> temperate nor<strong>the</strong>rn hemisphere. It also occurs scattered inland in England on3 -++-___...-..L+_road verges, presumably spreading with <strong>the</strong> use <strong>of</strong> de-icing salt. What is surprising is thatnei<strong>the</strong>r Danish scurvy-grass (Cochlearia danica) nor saltmarsh grass (Puccinellia distans) havebeen found on <strong>the</strong> <strong>Forest</strong> verges although we have looked for <strong>the</strong>m; <strong>the</strong>y are classic roadsidesalt plants which are now widespread on verges.4 5Spergularia rubr8. Sand-spurrey.<strong>Ashdown</strong> <strong>Forest</strong>, 1895, T. Hilton IBTN}.We only have three records; on cinders in car park, Isle <strong>of</strong> Thorns (418.303), 1995, <strong>Flora</strong>meeting; track in Streeter's Rough 142.291, 1993, TR; Black Hill area 147.31}, 1994, SBRS.This is yet ano<strong>the</strong>r annual which is more frequent on light, freely-drained sands, and it3 -f-.,......-----l...j- may not grow well on <strong>the</strong> siltier soils <strong>of</strong> <strong>Ashdown</strong> <strong>Forest</strong>.It is scattered through Britain and decreasing (Rich & Woodruff 1996). It occurs inEurope, Asia, North Africa and North America.4 5
58Lychnis flos-cuculi. Ragged Robin, Ragged Jack, Bachelor's buttons.Common on <strong>the</strong> <strong>Forest</strong> (Hall 1980).Usually found in damp or wet places in meadows, heathy grassland, on clayey rides andin open patches in woodland. In deep shade it flowers sparsely or not at all, and <strong>the</strong> hairless,narrow leaves are <strong>of</strong>ten a puzzle when found vegetatively.3 Widespread in Britain and Europe, but decreasing in England (Rich & Woodruff 1996).4 5*Lychnis chalcedonica. Maltese-cross.One clump with four fruiting stems just outside Lavender Platt (402.332), 1995, PW.This species is widely grown in gardens and occurs as a persistent escape elsewhere inBritain. It is native in eastern Europe.3 -++-----'-+-4 5Si/ene vulgaris. Bladder campion, Snappers.Tetrad 43F (Hall 1980).Entrance to Spring Garden (46.27), 1995, B. Hoath & A. Main, and apparently presentfor a long time before that.In Sussex widespread on <strong>the</strong> chalk and sometimes introduced with soils elsewhere,3 -++-----'-+__ possibly <strong>the</strong> origin <strong>of</strong> Hall's record. Widespread in Britain and Europe, but decreasing inEngland (Rich & Woodruff 1996). Also found in temperate Asia and North Africa.4 55i1ene latifolia (5. alba). White campion.Tetrad 42N (Hall 1980).We have three records, confirming its scarcity: Cold harbour Manor, bank by road andprobably introduced (399.326), 1995, TR; introduced with soil and mown, Goat car park(401.326), 1987-1995, PW; lane south <strong>of</strong> Nutley (44.26), 1993, MM. It is usually an annual3 --++_-----'4- <strong>of</strong> disturbed, open places such as arable field margins, but <strong>the</strong> plant at Goat car park seems to4 5be perennial perhaps due to mowing.Very common on <strong>the</strong> chalk in Sussex and occasional elsewhere (Hall 1980), but notreally a plant <strong>of</strong> <strong>the</strong> <strong>Forest</strong>. Widespread in Britain, Europe, western Asia and North Africa.5ilene dioica. Red campion, Bachelor's buttons.Common on <strong>the</strong> <strong>Forest</strong> (Hall 1980).Found mainly around <strong>the</strong> edge <strong>of</strong> <strong>the</strong> <strong>Forest</strong>, usually in nitrogen-rich, shaded places suchas river alluvium, old woodland edges and hedges. Usually <strong>the</strong> populations are small, but it isabundant on verge east <strong>of</strong> Fairwarp (473.266).It is relatively susceptible to frost damage, drought and waterlogging, and tends to occurin sheltered, open woodland (Baker 1947). Widespread in Britain and western Europe as wellas central Asia, North Africa and Greenland.We have not recorded any <strong>of</strong> <strong>the</strong> S. dioica x latifolia hybrid with pink flowers which is4 5usually quite widespread where <strong>the</strong> parents meet, probably because S. /atifo/ia is souncommon on <strong>the</strong> <strong>Forest</strong>. The two parent species are almost completely inter-fertile, and whilst S. latifolia is usuallypollinated by moths at night and S. dioica by bees during <strong>the</strong> day, <strong>the</strong>re are enough o<strong>the</strong>r pollinators around to ensureregular gene flow between <strong>the</strong>m; it seems that <strong>the</strong> species are maintained by <strong>the</strong> ecological differences, <strong>the</strong> hybrids beingnei<strong>the</strong>r properly suited to woods or fields (D. Goulson).
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Flora of Ashdown Forest,-,;I
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Copyright Tim Rich, Pat Donovan, Pa
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ACKNOWLEDGEMENTSWe would like to th
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THE HABITATS OF ASH DOWN FOREST3His
- Page 12 and 13: Near the bottom of all the non~wood
- Page 14 and 15: Conopodium majusCrataegus laevigata
- Page 16 and 17: Fincham (1995) published a small ph
- Page 18 and 19: 13ANAL VS IS OF THE FLORA DATAIn th
- Page 20 and 21: 15The number of different visits to
- Page 22 and 23: 17Hedgerows Disturbed/waste ground
- Page 24 and 25: 19The frequencies with which specie
- Page 26 and 27: 21(r~0.434, pO.l).-0.059,•300t350
- Page 28 and 29: 23A second type of comparison has b
- Page 30 and 31: 25Similarly, environmental variable
- Page 32 and 33: SPECIES ACCOUNTS27Lat'1n names foll
- Page 34 and 35: 29EQUISETACEAEEquisetum f/uviatile.
- Page 36 and 37: One plant on a vertical, north-faci
- Page 38 and 39: Viola pa/ustris in an open alder co
- Page 40 and 41: 35DRYOPTERIDACEAEPolystichum aculea
- Page 42 and 43: 37BLECHNACEAEBlechnum spicant. Hard
- Page 44 and 45: 39AngiospermsLAURACEAE* Laurus nobi
- Page 46 and 47: 41Ranunculus repens, Creeping butte
- Page 48 and 49: wet mud, small streams, ditches and
- Page 50 and 51: 45*Meconopsis cambrica. Welsh poppy
- Page 52 and 53: 47Ulmusx hollandica f?U. glabra x m
- Page 54 and 55: 49FAGACEAEFagus sylvatica. Beech.Fo
- Page 56 and 57: A special large oak at Chuck Hatch,
- Page 58 and 59: Cory/us avel/ana. Hazel.Recorded in
- Page 60 and 61: plants in the autumn; we have not r
- Page 64 and 65: 59*Saponaria officinalis. Soapwort,
- Page 66 and 67: 61Dolygonum arenastrum. Equal-leave
- Page 68 and 69: 63Rumex conglomeratus. Clustered do
- Page 70 and 71: 65Hypericum humifusum. Trailing S1
- Page 72 and 73: 67DROSERACEAEDrosera rotundifolia.
- Page 74 and 75: 69Viola laetea. Pale dog-violet.Thi
- Page 76 and 77: 71* Salix alba. White willow.3 -+-t
- Page 78 and 79: 73Salix repens var. repens (var. er
- Page 80 and 81: 75*Rorippa sylvestris. Creeping yel
- Page 82 and 83: 77* Thlaspi arvense. Field pennycre
- Page 84 and 85: 79aphanus raphanistrum. Wild radish
- Page 86 and 87: ica tetralix. Cross~leaved heath.As
- Page 88 and 89: centre of the Forest on alluvial so
- Page 90 and 91: It is predominantly a plant which o
- Page 92 and 93: 87SAXIFRAGACEAE[*Darmera pe/tata. I
- Page 94 and 95: 89insipid in flavour IHaskel1 1960)
- Page 96 and 97: 91ection Rubus. Subsection Hiemales
- Page 98 and 99: 93Rubus pr%ngatus.Nutley (45.27), 1
- Page 100 and 101: 95bus vestitus.Newbridge, 1969, B.
- Page 102 and 103: 97Rubus largificus.3 -+t--------'+_
- Page 104 and 105: Potentilla anserina. Silverweed.Com
- Page 106 and 107: 101Geum urbanum. Wood avens.Primari
- Page 108 and 109: 103osa x pseudorusticana fR. arvens
- Page 110 and 111: 105Prunus spinosa. Blackthorn, Wint
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orbus aucuparia. Rowan, Mountain as
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109r*Crataegus persimilis. Broad-le
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111icia tetrasperma. Smooth tare.Fo
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113Trifolium rep ens. White clover"
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115Genista tinctoria. Dyer's greenw
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*Genista anglica. Petty whin, Needl
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Myriophyllum alterniflorum. Alterna
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121Epilobium montanum x obscurum.Fo
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habitats.It lives up to the name of
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125,UPHORBIACEAEVlercurialis perenn
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Although one of Britain's tiniest p
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129*Oxalis articulata. Pink-sorrel.
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*Erodium cicutarium. Common stork's
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Widespread in Europe from the shore
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* Sison amomum. Stone parsley.Tetra
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137Centaurium pu/chellum. Lesser ce
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139CONVOLVULACEAEConvolvulus arvens
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141*Nymphoides pe/tata. Fringed wat
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Myosotis secunda (M. repens). Creep
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145Stachys 8lvensis. Field woundwor
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5cutellaria x hybrida (5. galericul
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Origanum vu/gare. Marjoram.Near Fis
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CALLlTRICHACEAE151Ca/litriche is a
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153Fraxinus excelsior. Ash.4 5Local
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~'Cymbalaria mural is. Ivy-leaved t
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157Veronica scutellata. Marsh speed
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159Sibthorpia europaea. Cornish mon
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161OROBANCHACEAEOrobanche rapum~gen
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163RUBIACEAESherardia arvensis. Mad
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165Viburnum OpUIU5.Guelder-rose, St
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167Valeriana officina/is 5.1. Commo
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169Cirsium vu/gate. Spear thistle.W
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171We suspect that Leontodon hispid
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173Mycelis muralis. Wall lettuce.Se
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175Taraxacum exceJlens.Scattered al
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177Taraxacum sinuatum.Recorded from
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179Crepis capillaris. Smooth hawk'
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181Hieracium. Hawkweeds.3134 5Hiera
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*So/idago gigantea. Early goldenrod
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ecorded with Chamaemelum are Matric
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*Senecio x albescens (S. cineraria
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*Petasites japonicus. Giant butterb
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*E1odea canadensis. Canadian waterw
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Arum maculatum. Lords~and~ladies, C
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Juncus x surrejanus (J. acutiflorus
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fences or topography. It may domina
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Eleocharis multicaulis. Many-stalke
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A hybrid first described by Col em
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203Carex riparia. Greater pond~sedg
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Material which initially looked lik
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207Festuca pratensis. Meadow fescue
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209Cynosurus cristatus. Crested dog
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211Glyceria f/uitans. Floating swee
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213Deschampsia cespitosa. Tufted ha
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Agrostis gigantea. Black bent.Tetra
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217Bromus lepidus. Slender soft~bro
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* Triticum aestivum. Bread wheat.A
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221L1L1ACEAENarthecium ossifragum.
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223*Allium triquetrum. Three-Gornar
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Trumpet length(mm)454035302520•
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227ORCHIDACEAEEpipactis purpurata.
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een unable to refind any bog orchid
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231DactylorhizB praetermissa. South
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Rachel A. Nicholson (1934-). Rachel
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235REFERENCESAbbas, M. H. (1979). S
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Graham, G. G. & Primavesi, A. L. (1
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Rich, T. C. G. & Smith, P. A. (1996
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GAZETTEER241This gazetteer has been
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Smockfarthing 40.31Smuggler's car p
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Asple adi+cuttr int~~~tl·Bteel SI!
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Broad bean 111 Cardamine x hausskne
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Fen bedstraw 163 Fumitory 46 Greate
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Leucanthemum maximum 186 Many-flowe
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Quercus ilex 50 Rorippa x ster/lis
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Sticky mouse-ear 56 Thale cress 73