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Flora of Ashdown Forest - Botanical Society of the British Isles

Flora of Ashdown Forest - Botanical Society of the British Isles

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146Common throughout most <strong>of</strong> south-eastern Britain but becoming rarer in <strong>the</strong> west and restricted to nutrientrichsites. Widespread over much <strong>of</strong> Europe except <strong>the</strong> far north and south. Also found in Himalaya and Japan.Lamium purpureum. Red deadnettle.This species was recorded in 90% <strong>of</strong> <strong>the</strong> tetrads in Sussex by Hall (1980) and isextremely common all over sou<strong>the</strong>rn England. It is, however, present in only 30% <strong>of</strong><strong>the</strong> 1-km squares in our area. Its sites on <strong>the</strong> <strong>Forest</strong> are strongly associated withhuman activity, occurring in gardens, disturbed areas on road verges and track sidesor as a pavement weed in <strong>the</strong> villages. It is absent from large areas <strong>of</strong> woodland andheathland.Very common throughout Britain and much <strong>of</strong> Europe.4 5Ga/eopsis tetrahit s./. Hemp~nettles.This aggregate is composed <strong>of</strong> G. tetrahit sensu stricto and G. bifida. They can only be distinguished in flower soour vegetative plants have been mapped as <strong>the</strong> aggregate. The aggregate is shown as common in Hall (1980) but<strong>the</strong>re is little reliable indication <strong>of</strong> <strong>the</strong> relative frequencies <strong>of</strong> <strong>the</strong> two segregates.Plants <strong>of</strong> <strong>the</strong> aggregate generally occur in disturbed habitats such as paths ides, <strong>the</strong> edges <strong>of</strong> arable fields andheaths, in light secondary woodland and by hedges. They are usually summer annuals and flower late in <strong>the</strong>season.Ga/eopsis tetrahit s.s. Common hemp~nettle.Confirmed in only three localities around <strong>the</strong> edge <strong>of</strong> <strong>the</strong> <strong>Forest</strong> during our survey.Common throughout Britain and Europe except <strong>the</strong> far south-east.Ga/eopsis bifida. Bifid hemp~nettle.Found to be more common than G. tetrahit during our survey despite <strong>the</strong> fact that Hall (1980) has no records for<strong>the</strong> area. This is certainly due to under-recording <strong>of</strong> this species in <strong>the</strong> past. The national distribution is not clearbut it is certainly present over much <strong>of</strong> <strong>the</strong> range <strong>of</strong> G. tetrahit s.s.In Finland, studies have shown that 96-100% <strong>of</strong> <strong>the</strong> seeds are dormant for at least ten years (Hintikka 1987).Seeds are probably stimulated to germinate by clear-felling and burning, and plants can dominate areas afterforestry work has been carried out. This strategy enables it to colonise disturbed competition-free habitats, Itbehaves similarly on <strong>Ashdown</strong> <strong>Forest</strong>, occurring on small disturbed areas and retreating into <strong>the</strong> seed bank as <strong>the</strong>vegetation closes over again.Ga/eopsis tetrahit s./, Galeopsis tetrahit 5.5, Ga/eopsis bifida3 +~-..------"Bf- 3-++_---~+- 3-++----~-+-4 5 4 5 4 5ScuteJ/aria ga/ericulata. Skullcap.Bogs near Gills Lap, <strong>Ashdown</strong> <strong>Forest</strong> (Coleman 1836). <strong>Forest</strong> Row, 1910, Miss P.Stockdale (BEX). Occasional (Hall 1980).We have only accepted two modern records, but <strong>the</strong>re could be more: Old Mill(48.30), 1995, AK; stream woodland near Cowfield (49.31), 1995, TR & PA. In3 -++_----'"'-+_ 1993 vegetative plants with too<strong>the</strong>d leaves were recorded widely in woodland, rides,ditches and streams and ponds, but were withdrawn in 1994 after we had discoveredthat S. x hybrida was much more common. It is quite possible that some <strong>of</strong> <strong>the</strong>records in Hall may be for <strong>the</strong> hybrid (see below).4 5S. galer/culata is a plant <strong>of</strong> ra<strong>the</strong>r wet, swampy places, favouring <strong>the</strong> edges <strong>of</strong>streams and ponds or wet grassland. It is a lowland plant with a wide distribution, occurring throughout Europe,in Asia, Algeria and North America, In <strong>the</strong> UK it is widely distributed over most <strong>of</strong> <strong>the</strong> country except in north-eastEngland and eastern Scotland. It is decreasing in England (Rich & Woodruff 1996).

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