It is predominantly a plant which occurs near <strong>the</strong> coast in Britain, except for <strong>the</strong> New <strong>Forest</strong> and <strong>the</strong> Weald, but isscattered through lowland Europe, south~west Asia, North Africa and <strong>the</strong> Americas. It is <strong>of</strong>ten associated with Radiatalino/des which has a similar ecology, and both are decreasing in England (Rich & Woodruff 1996).GROSSULARIACEAE*Ribes rubrum. Red currant.Common on <strong>the</strong> <strong>Forest</strong> (Hall 1980),Usually found in woodland, <strong>of</strong>ten along stream banks 'In alder carr but also in scrub andhedges. Away from <strong>the</strong> valleys it is likely to have been spread from gardens by birds. It usedto be grown at <strong>the</strong> Wych Cross fruit farm.It is predominately western in distribution in Europe, but is widely introduced and it isvirtually impossible to establish its native range. It is increasing in England (Rich & Woodruff1996),Some garden cultivars may be R. rubrum x spicE/tum.85*Ribes nigrum. Black currant, Gazel.Crowborough Warren, 1899, E, Gilbert (TLS), Occasional on west side <strong>of</strong> <strong>the</strong> <strong>Forest</strong> (Hall1980),It is now quite widespread, <strong>of</strong>ten on wetter ground than <strong>the</strong> o<strong>the</strong>r Ribes species. Itusually occurs on wet, rich alluvial soils in woodlands as shown in <strong>the</strong> east <strong>of</strong> <strong>the</strong> <strong>Forest</strong>, butit also occurs on quite dry roads ides around Wych Cross where it may have been bird-sownfrom <strong>the</strong> former fruit farm. Deer do not eat it.In Sussex it occurs predominately on <strong>the</strong> Tunbridge Wells Sands and Lower Greensand. Itis mainly nor<strong>the</strong>rn in distribution in Europe, and is probably native in swampy woodland in4 5eastern Europe and Asia. It is increasing in England (Rich & Woodruff 1996).It can be picked out from a distance from R. rubrum by <strong>the</strong> more sharply, deeply lobed leaves; close up it has amberglands under <strong>the</strong> leaves (absent in o<strong>the</strong>r species). Rubbed leaves usually smell <strong>of</strong> black currant though oddly some blackfruitedbushes opposite Wych Cross Nursery do not.*Ribes sanguineum, Flowering currant.,---;--::==------;---, Scattered on roadsides and near habitation; one plant on roadside south <strong>of</strong> Nutley (448.267),1994, TR, ano<strong>the</strong>r east <strong>of</strong> Nutley (444,2771, 1994, PD & AK, and ano<strong>the</strong>r in Long car park(426.311), 1995, TR, and elsewhere.Widely grown in gardens, and occasionally spreading into <strong>the</strong> wild throughout much <strong>of</strong>3 -+-t-~----'-t- Britain, especially in <strong>the</strong> north and west, and is increasing in England (Rich & Woodruff 1996).It is native in western North America.Garden plants with larger yellowish-pinkish flowers may be R. x gordonianum which4 5could also occur.*Ribes uva-crispa. Gooseberry, Goosegog.Frequent on <strong>the</strong> <strong>Forest</strong> (Hall 1980),Our records match <strong>the</strong> pattern shown by Hall. It is widely scattered in hedges and youngwoodland around <strong>the</strong> <strong>Forest</strong>, <strong>of</strong>ten near habitation.Widespread in Britain and Europe, and increasing in England (Rich & Woodruff 1996); italso occurs in Morocco. The form which is probably native in western Europe has short,dense, patent non-glandular hairs on <strong>the</strong> berries. Plants with a mixture <strong>of</strong> long glandular hairsand short hairs, or those with glabrous berries, are likely to be <strong>of</strong> garden origin, Plants withonly glandular hairs on <strong>the</strong> berries occur in <strong>the</strong> Carpathians.The first edition <strong>of</strong> Clapham, Tutin & Warburg's <strong>Flora</strong> <strong>of</strong> <strong>the</strong> <strong>British</strong> <strong>Isles</strong> had a minorprinting error giving <strong>the</strong> size <strong>of</strong> <strong>the</strong> fruits as 10-20 cm instead <strong>of</strong> 10-20 mm; it was affectionately known as <strong>the</strong> giantgooseberry edition!CRASSULACEAE*Crassula helmsii. New Zealand pigmyweed, Australian swamp stonecrop.No botanist has a good word for this Australasian species which is currently spreading rapidly in Britain. It probably initiallyspread as a throw-out from aquaria and garden ponds, and subsequently under its own steam. It grows in a range <strong>of</strong> waterconditions from acidic to alkaline and even saline water, from 3 metres deep to 0.5 metres above <strong>the</strong> normal water level,and it seems to be virtually indestructible. Perhaps <strong>the</strong> first positive thing to be said is that a carpet <strong>of</strong> flowers in <strong>the</strong> pondat Tompset's Bank filled <strong>the</strong> air with a lovely scent <strong>of</strong> honey in June 1995.
86We have found it in ponds at Kidbrooke Park (418.339), 1995, TR & PA; Tompset'sBank (424.338), 1995, PW; Pippingford Park, introduced with o<strong>the</strong>r aquatics (448.299 and445.302), 1995, <strong>Flora</strong> meetings; established in a tiny stream in a meadow on slope east <strong>of</strong>Nutley Post Office (445.275), 1995, TR, a more unusual habitat.3 There were only two records for Sussex in Hall (1980), but Briggs (1990) added 13more, and <strong>the</strong>re are even more now, It is now also widespread and increasing in England(Rich & Woodruff 1996).4 5 Control measures were being investigated at Weir Wood reservoir in 1994, where <strong>the</strong>rewere vast areas lining <strong>the</strong> shore at <strong>the</strong> normal water line, Sheets <strong>of</strong> black plastic were pinnedover areas which probably killed <strong>the</strong> plants by heat ra<strong>the</strong>r than shade; <strong>the</strong> Crassula had recovered by 1995. O<strong>the</strong>r controlmeasures suggested seem to involve major herbicide application, and it has yet to be eliminated from any site where it hasbecome well-established. Ano<strong>the</strong>r alien, Hydrocoty/e ranunculoides, seems to be out-competing it at Piltdown Pond!It seems to be insensitive to <strong>the</strong> wea<strong>the</strong>r. It survived <strong>the</strong> drought in 1995 at Weir Wood, turning slightly brown butrecovered rapidly. The only o<strong>the</strong>r aquatic with it was Littorella unit/ora which seems able to put its leaves up through <strong>the</strong>cushions. It grows throughout <strong>the</strong> winter and does not seem to be frost sensitive, probably in part because it is protectedby <strong>the</strong> water.*Semperv;vum tectorum. House-leek, Sengreen, Sungreen, Welcome home husband however drunk you be.Nutley, R. A. Boniface (Wolley-Dod 1937). Not refound in Nutley in 1995, PR.Once frequent on <strong>the</strong> ro<strong>of</strong>s <strong>of</strong> houses where it was planted in <strong>the</strong> belief it was a protection against lightning, but nowrarely seen in Sussex (plants still survive on a ro<strong>of</strong> nearby at Balcombe). Native in sou<strong>the</strong>rn Europe where it occurs onrocky slopes in <strong>the</strong> lowlands and mountains, almost a habitat match for ro<strong>of</strong>s and walls.Sedum teleph;um. Orpine, Livelong.• •3 -l--I-----Lt-~-~------~ <strong>Ashdown</strong> <strong>Forest</strong>, M. R. Dixon, 1886 (Arnold 1887). Frequent on <strong>the</strong> <strong>Forest</strong> (Hall 1980).Mainly recorded around <strong>the</strong> edges <strong>of</strong> <strong>the</strong> <strong>Forest</strong>, typically on hedge banks. Pathside,Horncastle Wood (394.311), 1994, DB; <strong>Forest</strong> Row, old railway line (43.34), 1987-1995 andspreading, PW; abundant along drive near Hunters Farm (447.264), 1994, TR; pathside south<strong>of</strong> Water Farm (441.335), 1995, PW; path, <strong>Forest</strong> Row (443.348), 1995, TR & PA; 46.32,1993, TR & EL. We have not looked to see which subspecies occur - <strong>the</strong> characters seem towork better on dried material than in <strong>the</strong> field.45It does not seem to be eaten by rabbits - plants along <strong>the</strong> drive to Hunter's Farm wereuntouched despite still being green and fleshy whilst everything else had been nibbled to <strong>the</strong>ground during <strong>the</strong> 1995 drought. It is apparently eaten by sheep and goats but not by horses.It is scattered in Britain in <strong>the</strong> lowlands, and although it will survive at altitude its growth rate is strongly dependenton temperature (Woodward 1987). Widespread in Europe, temperate Asia and North America.*Sedum spurium. Caucasian stonecrop.Garden rubbish, <strong>Forest</strong> Row (434.343), 1994, TR & NM, not surviving to 1995 as morerubbish was dumped on top.Native in south-east Asia. Occasionally established in Britain usually on calcareous rocks,and increasing in England (Rich & Woodruff 1996).3 --l--I-----4-4 5*Sedum acre. Biting stonecrop, Poor man's pepper.Recorded in tetrad 42N in Hall (1980), presumably from Nutley (perhaps once on a ro<strong>of</strong> with <strong>the</strong> Sempervivum) and notrefound. An interesting species which is mainly a calcicole occurring on soils above c. pH 6.0, but quite widely recordedon walls and calcareous sands. It is also grown in gardens.It is widespread in Britain. It occurs widely in Eurasia and North Africa.*Sedum album. White stonecrop.All our records are <strong>of</strong> plants escaping from gardens or introduced with soil. Naturalized onrailway bridge, <strong>Forest</strong> Row (438.347), 1994, TR & NM; introduced with soil by pond,Pippingford Park (44.30), 1995, PO & RN; spreading on graves, Fairwarp Church (466.268),1995, TR; Crow borough Training camp, possibly on bare ground near <strong>of</strong>fices (49.29), 1995,3 RN.Possibly native on limestones in <strong>the</strong> south-west <strong>of</strong> Britain, but not native in Sussex whereit mainly occurs on walls near <strong>the</strong> sea (Hall 1980). Increasing in England (Rich & Woodruff1996). Native in nor<strong>the</strong>rn and western Europe and North Africa.4 5
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Flora of Ashdown Forest,-,;I
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Copyright Tim Rich, Pat Donovan, Pa
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ACKNOWLEDGEMENTSWe would like to th
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THE HABITATS OF ASH DOWN FOREST3His
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Near the bottom of all the non~wood
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Conopodium majusCrataegus laevigata
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Fincham (1995) published a small ph
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13ANAL VS IS OF THE FLORA DATAIn th
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15The number of different visits to
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17Hedgerows Disturbed/waste ground
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19The frequencies with which specie
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21(r~0.434, pO.l).-0.059,•300t350
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23A second type of comparison has b
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25Similarly, environmental variable
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SPECIES ACCOUNTS27Lat'1n names foll
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29EQUISETACEAEEquisetum f/uviatile.
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One plant on a vertical, north-faci
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Viola pa/ustris in an open alder co
- Page 40 and 41: 35DRYOPTERIDACEAEPolystichum aculea
- Page 42 and 43: 37BLECHNACEAEBlechnum spicant. Hard
- Page 44 and 45: 39AngiospermsLAURACEAE* Laurus nobi
- Page 46 and 47: 41Ranunculus repens, Creeping butte
- Page 48 and 49: wet mud, small streams, ditches and
- Page 50 and 51: 45*Meconopsis cambrica. Welsh poppy
- Page 52 and 53: 47Ulmusx hollandica f?U. glabra x m
- Page 54 and 55: 49FAGACEAEFagus sylvatica. Beech.Fo
- Page 56 and 57: A special large oak at Chuck Hatch,
- Page 58 and 59: Cory/us avel/ana. Hazel.Recorded in
- Page 60 and 61: plants in the autumn; we have not r
- Page 62 and 63: Sagina procumbens. Procumbent pearl
- Page 64 and 65: 59*Saponaria officinalis. Soapwort,
- Page 66 and 67: 61Dolygonum arenastrum. Equal-leave
- Page 68 and 69: 63Rumex conglomeratus. Clustered do
- Page 70 and 71: 65Hypericum humifusum. Trailing S1
- Page 72 and 73: 67DROSERACEAEDrosera rotundifolia.
- Page 74 and 75: 69Viola laetea. Pale dog-violet.Thi
- Page 76 and 77: 71* Salix alba. White willow.3 -+-t
- Page 78 and 79: 73Salix repens var. repens (var. er
- Page 80 and 81: 75*Rorippa sylvestris. Creeping yel
- Page 82 and 83: 77* Thlaspi arvense. Field pennycre
- Page 84 and 85: 79aphanus raphanistrum. Wild radish
- Page 86 and 87: ica tetralix. Cross~leaved heath.As
- Page 88 and 89: centre of the Forest on alluvial so
- Page 92 and 93: 87SAXIFRAGACEAE[*Darmera pe/tata. I
- Page 94 and 95: 89insipid in flavour IHaskel1 1960)
- Page 96 and 97: 91ection Rubus. Subsection Hiemales
- Page 98 and 99: 93Rubus pr%ngatus.Nutley (45.27), 1
- Page 100 and 101: 95bus vestitus.Newbridge, 1969, B.
- Page 102 and 103: 97Rubus largificus.3 -+t--------'+_
- Page 104 and 105: Potentilla anserina. Silverweed.Com
- Page 106 and 107: 101Geum urbanum. Wood avens.Primari
- Page 108 and 109: 103osa x pseudorusticana fR. arvens
- Page 110 and 111: 105Prunus spinosa. Blackthorn, Wint
- Page 112 and 113: orbus aucuparia. Rowan, Mountain as
- Page 114 and 115: 109r*Crataegus persimilis. Broad-le
- Page 116 and 117: 111icia tetrasperma. Smooth tare.Fo
- Page 118 and 119: 113Trifolium rep ens. White clover"
- Page 120 and 121: 115Genista tinctoria. Dyer's greenw
- Page 122 and 123: *Genista anglica. Petty whin, Needl
- Page 124 and 125: Myriophyllum alterniflorum. Alterna
- Page 126 and 127: 121Epilobium montanum x obscurum.Fo
- Page 128 and 129: habitats.It lives up to the name of
- Page 130 and 131: 125,UPHORBIACEAEVlercurialis perenn
- Page 132 and 133: Although one of Britain's tiniest p
- Page 134 and 135: 129*Oxalis articulata. Pink-sorrel.
- Page 136 and 137: *Erodium cicutarium. Common stork's
- Page 138 and 139: Widespread in Europe from the shore
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* Sison amomum. Stone parsley.Tetra
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137Centaurium pu/chellum. Lesser ce
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139CONVOLVULACEAEConvolvulus arvens
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141*Nymphoides pe/tata. Fringed wat
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Myosotis secunda (M. repens). Creep
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145Stachys 8lvensis. Field woundwor
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5cutellaria x hybrida (5. galericul
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Origanum vu/gare. Marjoram.Near Fis
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CALLlTRICHACEAE151Ca/litriche is a
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153Fraxinus excelsior. Ash.4 5Local
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~'Cymbalaria mural is. Ivy-leaved t
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157Veronica scutellata. Marsh speed
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159Sibthorpia europaea. Cornish mon
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161OROBANCHACEAEOrobanche rapum~gen
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163RUBIACEAESherardia arvensis. Mad
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165Viburnum OpUIU5.Guelder-rose, St
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167Valeriana officina/is 5.1. Commo
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169Cirsium vu/gate. Spear thistle.W
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171We suspect that Leontodon hispid
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173Mycelis muralis. Wall lettuce.Se
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175Taraxacum exceJlens.Scattered al
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177Taraxacum sinuatum.Recorded from
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179Crepis capillaris. Smooth hawk'
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181Hieracium. Hawkweeds.3134 5Hiera
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*So/idago gigantea. Early goldenrod
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ecorded with Chamaemelum are Matric
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*Senecio x albescens (S. cineraria
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*Petasites japonicus. Giant butterb
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*E1odea canadensis. Canadian waterw
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Arum maculatum. Lords~and~ladies, C
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Juncus x surrejanus (J. acutiflorus
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fences or topography. It may domina
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Eleocharis multicaulis. Many-stalke
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A hybrid first described by Col em
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203Carex riparia. Greater pond~sedg
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Material which initially looked lik
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207Festuca pratensis. Meadow fescue
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209Cynosurus cristatus. Crested dog
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211Glyceria f/uitans. Floating swee
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213Deschampsia cespitosa. Tufted ha
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Agrostis gigantea. Black bent.Tetra
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217Bromus lepidus. Slender soft~bro
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* Triticum aestivum. Bread wheat.A
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221L1L1ACEAENarthecium ossifragum.
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223*Allium triquetrum. Three-Gornar
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Trumpet length(mm)454035302520•
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227ORCHIDACEAEEpipactis purpurata.
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een unable to refind any bog orchid
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231DactylorhizB praetermissa. South
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Rachel A. Nicholson (1934-). Rachel
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235REFERENCESAbbas, M. H. (1979). S
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Graham, G. G. & Primavesi, A. L. (1
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Rich, T. C. G. & Smith, P. A. (1996
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GAZETTEER241This gazetteer has been
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Smockfarthing 40.31Smuggler's car p
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Asple adi+cuttr int~~~tl·Bteel SI!
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Broad bean 111 Cardamine x hausskne
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Fen bedstraw 163 Fumitory 46 Greate
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Leucanthemum maximum 186 Many-flowe
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Quercus ilex 50 Rorippa x ster/lis
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Sticky mouse-ear 56 Thale cress 73