29EQUISETACEAEEquisetum f/uviatile. Water horsetail.Six tetrads (Hall 1980).Recorded frequently from <strong>the</strong> wetter valleys south <strong>of</strong> <strong>Forest</strong> Row, at Old Lodge and in apond in <strong>the</strong> corner <strong>of</strong> Furnace Wood. Almost all localities are in ponds, <strong>of</strong>ten where <strong>the</strong>re iso<strong>the</strong>r vegetation and bottoms choked with silt, or in gentle flushes in some <strong>of</strong> <strong>the</strong> springs.3 Many <strong>of</strong> <strong>the</strong> sites are shaded. It has been suggested to be particularly abundant in marl pitsbut this has not been confirmed.It is widespread in Britain from <strong>the</strong> rich ditch systems <strong>of</strong> Sussex grazing marshes such asPevensey Levels, to clear nutrient-poor lakes in Scotland. Jermy et al. (1978) note <strong>the</strong> ecology4 5is little understood. It is widespread in Europe, temperate Asia and North America.Equisetum x litorale (E. fluviatile x palustre). Shore horsetail.<strong>Ashdown</strong> <strong>Forest</strong>, c. 1955, R. A. Boniface (BRC). Not refound.Jermy et al. (1978) note that this hybrid is surprisingly frequent in Britain and Ireland given <strong>the</strong> dissimilarity <strong>of</strong> <strong>the</strong>habitats <strong>of</strong> <strong>the</strong> parents. There is a distinct cluster <strong>of</strong> records in <strong>the</strong> north Weald which is probably due to <strong>the</strong> efforts <strong>of</strong> asingle recorder.Equisetum arvense. Field horsetail.Common in tetrads around <strong>the</strong> edges <strong>of</strong> <strong>the</strong> <strong>Forest</strong> (Hall 1980).We have recorded 'It quite widely in arable fields, gardens, waste ground around <strong>the</strong> edges<strong>of</strong> <strong>the</strong> <strong>Flora</strong> area, and on roadsides on <strong>the</strong> <strong>Forest</strong>. It is distinctly uncommon on <strong>the</strong> <strong>Forest</strong>.Common and widespread in Britain, and recorded in Europe, central China, North America3 and Greenland.4 5Equisetum palustre. Marsh horsetail.Tetrads 33V, 43F and 431< (Hall 1980).Recorded from scattered squares around <strong>the</strong> edge <strong>of</strong> <strong>the</strong> <strong>Forest</strong>, and in <strong>the</strong> Chelwoodarea. Jermy et al. (1978) note that it requires moving ground water with a medium basecontent(calcium/magnesium), and it may occur where <strong>the</strong> soils are less acidic.3 -+-l---....-------Y- It is frequent in Sussex, and common and widespread in Britain, Europe, temperate Asiaand North America.4 5Equisetum telmateia. Great horsetail.Recorded from tetrads 42N, 43K, 43L, 43M and 43V in Hall (1980), some <strong>of</strong> which may have been from outside our area,but oddly not recorded at all during our survey. This species <strong>of</strong>ten occurs on wet clays in Sussex <strong>of</strong>ten by spring lineswhere <strong>the</strong>re is some water seepage, a habitat not present in most <strong>of</strong> Hall's tetrads on <strong>the</strong> <strong>Forest</strong>. It is found locallythroughout Europe, Asia, North Africa and North America.OPHIOGLOSSACEAEOphioglossum vulgatum. Adder's-tongue.Tetrads 43G (Hall 1980) and 42T (Briggs 1990), and refound in both.The main habitat on <strong>Ashdown</strong> <strong>Forest</strong> is on <strong>the</strong> more floristically-rich verges, clearly where<strong>the</strong>re is some base-enrichment from <strong>the</strong> road, and some patches may be 5 metres long.Twyford Lane (404.314), 1995, CM; near Goat cross roads (402.325), 1993, TR & NM; Priory3 Road (405.328), c. 1993, CM; both sides <strong>of</strong> Hindleap Road (418.319), 1995, CM; A275 nearWych Cross Place (419.315), 1990, CM; thousands on private meadows and lawns, OldCherry Cottage (41.33 and 42.33), 1995, PS; Chelwood Vachery drive (435.304), 1993, TR &4 5 NM; east <strong>of</strong> <strong>the</strong> Visitor Centre (434.323), 1992, NM and 1995, CM; verge 30 metres NE <strong>of</strong>car park, Gills Lap (468.315),1995, TR; verge 100 metres west <strong>of</strong> Greenwood Gate(478.314), 1995, TR; Poundgate verge (482.285), 1995, CM, big colony, 1995, PS; two fronds, Toll Lane wood(461.263), 1993, TR & NM, not refound SUbsequently.This small, grass-green fern is very difficult to see, and is probably generally under-recorded (e.g. Briggs 1990 gaveabout twice as many records for Sussex as HaJl 1980). It is found most easily in April or May before <strong>the</strong> vegetation closesover, and <strong>the</strong> fronds have usually died down by <strong>the</strong> end <strong>of</strong> June or been mown.
30It is widespread but rarely common in England, but rarer in Wales and Scotland where it is predominantly coastal(Jermy et al. 1978), and it is also widespread in Europe, Asia, North Africa and North America.Botrychium lunaria. Moonwort.On <strong>the</strong> <strong>Forest</strong> between Fisher's Gate and Maresfield, on <strong>the</strong> open part <strong>of</strong> <strong>the</strong> <strong>Forest</strong> between Crowborough Warren and <strong>the</strong>new church, and between Coleman's Hatch and <strong>Forest</strong> Row, E. Jenner (Arnold 1887). These are <strong>the</strong> last records for <strong>the</strong><strong>Forest</strong> where it is probably long extinct.The records <strong>of</strong> Botrychium from acid heathland are initially somewhat surprising, as it is usually a calcicole <strong>of</strong> welldrainedsites in rich mountain grassland, limestone grassland and sand dunes. However, it does occur on calcareousheaths in <strong>the</strong> Breckland, and on acidic soils such as Witley Common in Surrey with hea<strong>the</strong>r, and also under bracken(Lousley 1976). It has certainly declined in south~east England and was last recorded in Sussex near Brighton in 1965 (Hall1980). Not seen in Kent since 1947, and only in three squares in Surrey recently.Widespread in nor<strong>the</strong>rn Europe, and in <strong>the</strong> mountains in <strong>the</strong> south, and in Asia Minor, Himalaya and Australasia.OSMUNDACEAEOsmunda regalis. Royal fern.On <strong>Ashdown</strong> <strong>Forest</strong> (Forster 1816), )n some <strong>of</strong> <strong>the</strong> bogs on <strong>Ashdown</strong> <strong>Forest</strong> (Co)eman 1836),In <strong>the</strong> forests, rare, E. Jenner (Arnold 1887). <strong>Ashdown</strong> <strong>Forest</strong> but victim <strong>of</strong> vandalism (Pickard1930), Chuck Hatch, 1931, E, C, Wallace (Wolley-Dod 1937), Che(wood Gate (probablY42.30), two big clumps and several o<strong>the</strong>rs scattered around, 1948, 1949, G. Dent (Dent 1928-3+H(-!""---'-+-1953), Chelwood Gate 1955, R, A, Boniface (BRC),Native plants occur in <strong>the</strong> following sites: on <strong>the</strong> side <strong>of</strong> a slit trench (422.308) and in adrainage gully (c, 423,305) east <strong>of</strong> <strong>the</strong> Isle <strong>of</strong> Thorns, 1991 +, CM & NM; one plant in a root4 5 plate by <strong>the</strong> path near Hoadley's Farm (506,324), 1994, TR et ai, which disappeared when <strong>the</strong>root plate fell on it during <strong>the</strong> winter before it could produce spores; Broadstone Warren(42.33), CM, 1995. This last site is superb, with four large fertile clumps (one with fronds to 2 metres tall, and ano<strong>the</strong>repiphytic on <strong>the</strong> bole <strong>of</strong> an alder coppice) and about seven young plants in an open, flushed alder wood with beech fern.It has been taken frequently from <strong>the</strong> wild into gardens, and has probably re-established itself in <strong>the</strong> wild again fromspores. We have recorded it introduced in <strong>the</strong> following sites: planted in Chelwood Vachery around <strong>the</strong> bog pool(431,294), 1994, MR, PR & TR; one plant by a pond at Home Farm, Pippingford Park (443,304), 1993, PD & TR; severalclumps by a swimming pond, Northbank Wood (434,319), 1995, TR & PA where G, Dent regularly recorded it (Dent 1928-1953); planted around springs in Pheasantry (440.315 - not plotted) and present to at least 1986, M. Tebbutt; one bypond, Ridge Road (448,329), 1992 +, CM & NM; in <strong>the</strong> rockery at Oldlands Hall (476,275), 1995, PD et ai,Royal fern is one <strong>of</strong> those wonderful plants whose bright green foliage contrasts with <strong>the</strong> golden fertile fronds. In <strong>the</strong>Weald it tends to grow in valley bogs or damp woodland, and sometimes on <strong>the</strong> sand rocks, but is never now abundant,and Hall (1980) regards <strong>Ashdown</strong> <strong>Forest</strong> as <strong>the</strong> only native area in Sussex. Wolley-Dod (1937) gives 35 records but notesit was formerly more common, and Hall (1980) only gives half that number <strong>of</strong> tetrads including introduced sites.It is scattered through Britain mainly in areas with high rainfall, from sea cliffs in Cornwall to fens in East Anglia andbogs in <strong>the</strong> north and west, and even in a limestone crevice at an altitude <strong>of</strong> 400 metres on Ingleborough, Yorkshire. InIreland it is <strong>of</strong>ten abundant on abandoned peat cuttings. It is a western sub-Atlantic plant in Europe, and occurs widely in<strong>the</strong> tropics, and North America.MARSILEACEAEPilularia globulifera. Pillwort.In a bog at Quarry Brook [Quabrook = a boggy place] on <strong>the</strong> right <strong>of</strong> <strong>the</strong> road from Hartfield to <strong>Forest</strong> Row, E. Jenner(Arnold 1887); this record is from what must have been an excellent boggy site, but is now drained. <strong>Ashdown</strong> <strong>Forest</strong>near Nutley, 1957, R, A, Boniface (BRC),Pilu/aria is now probably extinct in Sussex. Wolley-Dod (1937) recorded 19 sites, and <strong>the</strong> sites at Horsham andPiltdown (Hall 1980) have gone. A more recent site near Crawley (Briggs 1990) was recently destroyed by landfill. It isnow a Nationally Scarce Species recorded in 162 10-km squares before 1970 compared to only 90 since 1970. Most <strong>of</strong><strong>the</strong> decline occurred between 1918 and 1950 (Jermy 1994).Pilu/aria is a plant <strong>of</strong> lowland nutrient-poor pond and lake margins where <strong>the</strong>re is some disturbance or bare ground. Itis a western European endemic, with <strong>the</strong> bulk <strong>of</strong> <strong>the</strong> sites occurring in lowland Britain, France, nor<strong>the</strong>rn Germany andSweden. It is declining in much <strong>of</strong> mainland Europe. The New <strong>Forest</strong> is thought to support <strong>the</strong> largest concentration <strong>of</strong>sites.HYMENOPHYLLACEAEHymenophyJ/um tunbrigense. Tunbridge filmy-fern.Furnace Wood, c. 1955, FR (tetrad 42T in Hall 1980), Thought lost due to tree felling but one patch refound 1975, M,Rickard, 1985, FR (also reported just outside <strong>the</strong> <strong>Flora</strong> area at Rock Wood on <strong>the</strong> opposite side <strong>of</strong> <strong>the</strong> valley to FurnaceWood, OS, but not seen recently).
- Page 1 and 2: Flora of Ashdown Forest,-,;I
- Page 3 and 4: Copyright Tim Rich, Pat Donovan, Pa
- Page 5: ACKNOWLEDGEMENTSWe would like to th
- Page 8 and 9: THE HABITATS OF ASH DOWN FOREST3His
- Page 12 and 13: Near the bottom of all the non~wood
- Page 14 and 15: Conopodium majusCrataegus laevigata
- Page 16 and 17: Fincham (1995) published a small ph
- Page 18 and 19: 13ANAL VS IS OF THE FLORA DATAIn th
- Page 20 and 21: 15The number of different visits to
- Page 22 and 23: 17Hedgerows Disturbed/waste ground
- Page 24 and 25: 19The frequencies with which specie
- Page 26 and 27: 21(r~0.434, pO.l).-0.059,•300t350
- Page 28 and 29: 23A second type of comparison has b
- Page 30 and 31: 25Similarly, environmental variable
- Page 32 and 33: SPECIES ACCOUNTS27Lat'1n names foll
- Page 36 and 37: One plant on a vertical, north-faci
- Page 38 and 39: Viola pa/ustris in an open alder co
- Page 40 and 41: 35DRYOPTERIDACEAEPolystichum aculea
- Page 42 and 43: 37BLECHNACEAEBlechnum spicant. Hard
- Page 44 and 45: 39AngiospermsLAURACEAE* Laurus nobi
- Page 46 and 47: 41Ranunculus repens, Creeping butte
- Page 48 and 49: wet mud, small streams, ditches and
- Page 50 and 51: 45*Meconopsis cambrica. Welsh poppy
- Page 52 and 53: 47Ulmusx hollandica f?U. glabra x m
- Page 54 and 55: 49FAGACEAEFagus sylvatica. Beech.Fo
- Page 56 and 57: A special large oak at Chuck Hatch,
- Page 58 and 59: Cory/us avel/ana. Hazel.Recorded in
- Page 60 and 61: plants in the autumn; we have not r
- Page 62 and 63: Sagina procumbens. Procumbent pearl
- Page 64 and 65: 59*Saponaria officinalis. Soapwort,
- Page 66 and 67: 61Dolygonum arenastrum. Equal-leave
- Page 68 and 69: 63Rumex conglomeratus. Clustered do
- Page 70 and 71: 65Hypericum humifusum. Trailing S1
- Page 72 and 73: 67DROSERACEAEDrosera rotundifolia.
- Page 74 and 75: 69Viola laetea. Pale dog-violet.Thi
- Page 76 and 77: 71* Salix alba. White willow.3 -+-t
- Page 78 and 79: 73Salix repens var. repens (var. er
- Page 80 and 81: 75*Rorippa sylvestris. Creeping yel
- Page 82 and 83: 77* Thlaspi arvense. Field pennycre
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79aphanus raphanistrum. Wild radish
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ica tetralix. Cross~leaved heath.As
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centre of the Forest on alluvial so
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It is predominantly a plant which o
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87SAXIFRAGACEAE[*Darmera pe/tata. I
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89insipid in flavour IHaskel1 1960)
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91ection Rubus. Subsection Hiemales
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93Rubus pr%ngatus.Nutley (45.27), 1
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95bus vestitus.Newbridge, 1969, B.
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97Rubus largificus.3 -+t--------'+_
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Potentilla anserina. Silverweed.Com
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101Geum urbanum. Wood avens.Primari
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103osa x pseudorusticana fR. arvens
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105Prunus spinosa. Blackthorn, Wint
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orbus aucuparia. Rowan, Mountain as
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109r*Crataegus persimilis. Broad-le
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111icia tetrasperma. Smooth tare.Fo
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113Trifolium rep ens. White clover"
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115Genista tinctoria. Dyer's greenw
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*Genista anglica. Petty whin, Needl
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Myriophyllum alterniflorum. Alterna
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121Epilobium montanum x obscurum.Fo
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habitats.It lives up to the name of
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125,UPHORBIACEAEVlercurialis perenn
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Although one of Britain's tiniest p
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129*Oxalis articulata. Pink-sorrel.
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*Erodium cicutarium. Common stork's
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Widespread in Europe from the shore
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* Sison amomum. Stone parsley.Tetra
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137Centaurium pu/chellum. Lesser ce
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139CONVOLVULACEAEConvolvulus arvens
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141*Nymphoides pe/tata. Fringed wat
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Myosotis secunda (M. repens). Creep
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145Stachys 8lvensis. Field woundwor
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5cutellaria x hybrida (5. galericul
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Origanum vu/gare. Marjoram.Near Fis
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CALLlTRICHACEAE151Ca/litriche is a
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153Fraxinus excelsior. Ash.4 5Local
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~'Cymbalaria mural is. Ivy-leaved t
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157Veronica scutellata. Marsh speed
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159Sibthorpia europaea. Cornish mon
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161OROBANCHACEAEOrobanche rapum~gen
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163RUBIACEAESherardia arvensis. Mad
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165Viburnum OpUIU5.Guelder-rose, St
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167Valeriana officina/is 5.1. Commo
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169Cirsium vu/gate. Spear thistle.W
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171We suspect that Leontodon hispid
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173Mycelis muralis. Wall lettuce.Se
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175Taraxacum exceJlens.Scattered al
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177Taraxacum sinuatum.Recorded from
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179Crepis capillaris. Smooth hawk'
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181Hieracium. Hawkweeds.3134 5Hiera
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*So/idago gigantea. Early goldenrod
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ecorded with Chamaemelum are Matric
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*Senecio x albescens (S. cineraria
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*Petasites japonicus. Giant butterb
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*E1odea canadensis. Canadian waterw
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Arum maculatum. Lords~and~ladies, C
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Juncus x surrejanus (J. acutiflorus
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fences or topography. It may domina
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Eleocharis multicaulis. Many-stalke
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A hybrid first described by Col em
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203Carex riparia. Greater pond~sedg
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Material which initially looked lik
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207Festuca pratensis. Meadow fescue
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209Cynosurus cristatus. Crested dog
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211Glyceria f/uitans. Floating swee
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213Deschampsia cespitosa. Tufted ha
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Agrostis gigantea. Black bent.Tetra
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217Bromus lepidus. Slender soft~bro
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* Triticum aestivum. Bread wheat.A
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221L1L1ACEAENarthecium ossifragum.
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223*Allium triquetrum. Three-Gornar
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Trumpet length(mm)454035302520•
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227ORCHIDACEAEEpipactis purpurata.
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een unable to refind any bog orchid
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231DactylorhizB praetermissa. South
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Rachel A. Nicholson (1934-). Rachel
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235REFERENCESAbbas, M. H. (1979). S
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Graham, G. G. & Primavesi, A. L. (1
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Rich, T. C. G. & Smith, P. A. (1996
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GAZETTEER241This gazetteer has been
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Smockfarthing 40.31Smuggler's car p
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Asple adi+cuttr int~~~tl·Bteel SI!
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Broad bean 111 Cardamine x hausskne
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Fen bedstraw 163 Fumitory 46 Greate
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Leucanthemum maximum 186 Many-flowe
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Quercus ilex 50 Rorippa x ster/lis
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Sticky mouse-ear 56 Thale cress 73