39AngiospermsLAURACEAE* Laurus nobilis. Bay.] -++-~~~-----'-+-Seedlings were found on a path and verge outside a garden near a bush at Cackle Street(452.266), 1995, RN & ER.This widely used pot herb is occasionally established in <strong>the</strong> wild in sheltered locations inBritain. As a native it occurs on <strong>the</strong> limestone along <strong>the</strong> Mediterranean coast and possibly <strong>the</strong>Atlantic coasts <strong>of</strong> Portugal and Spain, and is <strong>of</strong>ten now confined to inaccessible rocky cliffs.4 5NYMPHAEACEAE*Nymphaea alba. White water-lily.Cackle Street, 1933, G. Dent (Dent 1928-1953). Tetrads 42U and 43K (Hall 1980).Probably native in some localities in Sussex, but our records are probably always <strong>of</strong> liliesplanted in ponds and lakes. Sometimes <strong>the</strong> flowers are pink, as at Braberry Hatch and many]-++-.....'"--'"--L.j-o<strong>the</strong>r locations, and sometimes white as at Mill Wood, <strong>the</strong> pond by Ridge Road, and Ellison'sPond. During <strong>the</strong> 1995 drought some ponds dried out leaving whole plants exposed on drying4 5mud; <strong>the</strong>y seem to have survived quite well although a few were eaten by deer.The water in <strong>the</strong> ponds is <strong>of</strong>ten quite murky and dark, and <strong>the</strong> only o<strong>the</strong>r species to growwith it at all frequently is Potamogeton natans. The poor nutrient status <strong>of</strong> <strong>the</strong> ponds on <strong>the</strong>acidic soils probably also suits this species compared to Nuphar lutea, and indeed smallerNymphaea plants, sometimes distinguished as a separate subspecies, are widespread in <strong>the</strong> acidic poor waters <strong>of</strong> lochs andlochans in <strong>the</strong> north and west <strong>of</strong> Britain and Ireland.Widespread in Britain and Europe, though rare in <strong>the</strong> far north.Nuphar lutea. Yellow water-lily, Brandy bottle.<strong>Forest</strong> Row, Miss P. Stockdale (undated but c. 1910-1916; SEX). Still known in this area, withlarge patches possibly planted in a lake at <strong>Forest</strong> Row (421.347), 1995, TR, and probablynative as non-flowering submerged leaves in River Medway east <strong>of</strong> <strong>Forest</strong> Row (441.349),1995, TR & PA.] -++-~~~--'-+-This water-lily tends to occur in more nutrient-rich, calcareous waters than <strong>the</strong> whitewater-lily, and is <strong>of</strong>ten also found in deeper and running water. As well as <strong>the</strong> floating leavesit has submerged ones which are not best suited to muddy water, and its absence from <strong>the</strong>4 5<strong>Forest</strong> ponds may be due to <strong>the</strong>ir generally murky nutrient-poor water; it could also be absentbecause it has not been introduced.Common and widespread in Britain and Europe, though rare in <strong>the</strong> far north.CERATOPHYLLACEAECeratophyllum demersum. Rigid hornwort.Pond on <strong>the</strong> edge <strong>of</strong> <strong>the</strong> golf course at <strong>Forest</strong> Row (437.342), 1994; pond by Ridge Road(448.329), 1995; mucky pond by farm at Pricketts Hatch (443.270), 1995; north side <strong>of</strong>pond at <strong>Ashdown</strong> Park (431.320), 1995, all found by TR with a grapnel.This species floats freely in <strong>the</strong> water just below <strong>the</strong> surface, and can occur in open3 conditions or lurk under dense mats <strong>of</strong> duckweed, as at Prickets Hatch, where it seems to bequite shade-tolerant, an unusual characteristic for a water plant. It <strong>of</strong>ten occurs in nutrientrich45water, and is <strong>of</strong>ten abundant in arable areas where <strong>the</strong>re is high inorganic nitrogen in <strong>the</strong>water, probably from fertiliser run-<strong>of</strong>f (Goulder & Boatman 1971). The widespread increase ineutrophic water due to fertilisers may have resulted in an increase in this species in <strong>the</strong>lowlands.Occasional but probably under-recorded in Sussex (Briggs 1990 had about twice as many records as Hall 1980). Fewbotanists regularly carry a grapnel or are prepared to use it in smelly, dirty water, a potential habitat for both <strong>the</strong> hornwortand Weil's disease. Widespread in Britain, Europe, North Africa, Asia and North America,
40RANUNCULACEAECaltha palustris. Marsh-marigold, Kingcup.,---.. --===----.. , Recorded in six tetrads in Hall 11980),3 -++-. ... ----'_+_4 5We have it in seven 1-km squares most <strong>of</strong> which match Hall's except for two tetrads on<strong>the</strong> west side <strong>of</strong> <strong>the</strong> <strong>Forest</strong>, and also introduced at Chelwood Vachery around <strong>the</strong> spring1432,297), It is mainly found in <strong>the</strong> alder woods around <strong>the</strong> edges <strong>of</strong> <strong>the</strong> <strong>Forest</strong> le,g,Newbridge) but it is surprisingly rare; one might have imagined that it would be frequent in <strong>the</strong>wet, acidic flushes around <strong>the</strong> <strong>Forest</strong>.Widespread in <strong>the</strong> Weald, Britain and Europe, but decreasing in England (Rich & Woodruff1996). Also in temperate and arctic Asia, and North America.Anemone nemorosa. Wood anemone, Drops <strong>of</strong> snow.Quite widespread on <strong>the</strong> <strong>Forest</strong>, even in open situations on road verges and with brackenaway from woodland - <strong>the</strong> verges and banks on <strong>the</strong> 82026 around Fairwarp are lined withpatches, <strong>the</strong> white drifts most noticeable on sunny days in April. It may also be present in atleast some <strong>of</strong> <strong>the</strong> squares we have not had access to in <strong>the</strong> spring.Wolley-Dod (1937) notes that plants with lavender-co[oured flowers (var. caeru/ea DC.)were recorded by Talbot from <strong>Ashdown</strong> <strong>Forest</strong>. Occasional patches with blue-, pink- orpurple-f[ushed petals can now be found quite widely; blue-flowered plants were found by Goatcar park, RN (<strong>the</strong>se went purple when driedL PR reported one very striking patch atShepherd's car park with magenta petals coloured on both sides (var. purpurea DC.) amongstnormal white patches, and var. purpurea was also seen in woodland south <strong>of</strong> Londonderry Farm (446.289), 1982, ER.Plants with flowers <strong>of</strong> both colours occur at Maskett's Wood. As plants spread to form patches by rhizomes under <strong>the</strong>soil, sometimes different clones grow toge<strong>the</strong>r and can be distinguished by <strong>the</strong> slightly different shapes <strong>of</strong> leaves.Wood anemones are shade-tolerating ra<strong>the</strong>r than shade-demanding, and will grow quite successfully in <strong>the</strong> open. Theygrow best on moist to wet soils where <strong>the</strong> air humidity is high, and this is why <strong>the</strong>y are most frequently found in woodlandor shaded areas (Shirreffs 1985). Where <strong>the</strong> climate is dry <strong>the</strong>y are almost confined to woodlands (e.g. Lincolnshire), butwhere it is damper <strong>the</strong>y are less restricted (e.g. Derbyshire). They grow best on soils <strong>of</strong> a relatively heavy texture (e.g.clayey or silty), although <strong>the</strong>y will tolerate a range <strong>of</strong> soil conditions. The suitability <strong>of</strong> both <strong>the</strong> damp climate and <strong>the</strong> siltysoils may help explain <strong>the</strong>ir frequency in <strong>the</strong> open on Ashdawn <strong>Forest</strong>.Although <strong>the</strong>y contain a bitter poison, anemonin, which discourages grazing, <strong>the</strong>y may have decreased since <strong>the</strong>decline in grazing on <strong>the</strong> <strong>Forest</strong> due to <strong>the</strong> spread <strong>of</strong> bracken. They can tolerate light fires. The open conditions after <strong>the</strong>1987 storm caused some patches in woodland to flower better (AFN 15: 2-4).Common in Sussex (Hall 1980)' and widespread in Britain, Europe and western Asia.* Clematis vitalba. Traveller's-joy, Boy's bacca, Tom bacca, Old man, Old man's beard.Recorded in tetrad 33W and 43A in Hall 11980), but <strong>the</strong> latter is probably in 43B by Warrencar park where <strong>the</strong>re is an old conspicuous plant on <strong>the</strong> north side <strong>of</strong> <strong>the</strong> road.We have six records mainly from hedges and urban areas where it has escaped from3 -++------'_+_cultivation, for instance hedge at Horncastle 1390.324), 1995, TR; Warren car park 1416,320),pH 6.8, 1993, TR; scrambling over trees in young woodland, Tompset's Bank 1425,339),4 51995, PW; verge opposite Quarry car park 1470,319), 1993, PW.In Sussex it is common and widespread on <strong>the</strong> chalk, with a few records from <strong>the</strong> Weald(Hall 1980). It is widespread on chalky soils in lowland England, Europe, <strong>the</strong> Caucasus andNorth Africa.Ranunculus acris. Meadow buttercup, Crowfoot, Goldcup, Kingcup.Common on <strong>the</strong> <strong>Forest</strong> (Hall 1980),We have recorded it widely on <strong>the</strong> <strong>Forest</strong> on verges and in meadows, especially indamper areas, but it is rarely abundant. It avoids <strong>the</strong> more acidic soils. It appears to bereasonably resistant to herbicides but is <strong>of</strong>ten reduced in frequency where <strong>the</strong>y have beenapplied.It is widespread in Britain and temperate and arctic Eurasia.
- Page 1 and 2: Flora of Ashdown Forest,-,;I
- Page 3 and 4: Copyright Tim Rich, Pat Donovan, Pa
- Page 5: ACKNOWLEDGEMENTSWe would like to th
- Page 8 and 9: THE HABITATS OF ASH DOWN FOREST3His
- Page 12 and 13: Near the bottom of all the non~wood
- Page 14 and 15: Conopodium majusCrataegus laevigata
- Page 16 and 17: Fincham (1995) published a small ph
- Page 18 and 19: 13ANAL VS IS OF THE FLORA DATAIn th
- Page 20 and 21: 15The number of different visits to
- Page 22 and 23: 17Hedgerows Disturbed/waste ground
- Page 24 and 25: 19The frequencies with which specie
- Page 26 and 27: 21(r~0.434, pO.l).-0.059,•300t350
- Page 28 and 29: 23A second type of comparison has b
- Page 30 and 31: 25Similarly, environmental variable
- Page 32 and 33: SPECIES ACCOUNTS27Lat'1n names foll
- Page 34 and 35: 29EQUISETACEAEEquisetum f/uviatile.
- Page 36 and 37: One plant on a vertical, north-faci
- Page 38 and 39: Viola pa/ustris in an open alder co
- Page 40 and 41: 35DRYOPTERIDACEAEPolystichum aculea
- Page 42 and 43: 37BLECHNACEAEBlechnum spicant. Hard
- Page 46 and 47: 41Ranunculus repens, Creeping butte
- Page 48 and 49: wet mud, small streams, ditches and
- Page 50 and 51: 45*Meconopsis cambrica. Welsh poppy
- Page 52 and 53: 47Ulmusx hollandica f?U. glabra x m
- Page 54 and 55: 49FAGACEAEFagus sylvatica. Beech.Fo
- Page 56 and 57: A special large oak at Chuck Hatch,
- Page 58 and 59: Cory/us avel/ana. Hazel.Recorded in
- Page 60 and 61: plants in the autumn; we have not r
- Page 62 and 63: Sagina procumbens. Procumbent pearl
- Page 64 and 65: 59*Saponaria officinalis. Soapwort,
- Page 66 and 67: 61Dolygonum arenastrum. Equal-leave
- Page 68 and 69: 63Rumex conglomeratus. Clustered do
- Page 70 and 71: 65Hypericum humifusum. Trailing S1
- Page 72 and 73: 67DROSERACEAEDrosera rotundifolia.
- Page 74 and 75: 69Viola laetea. Pale dog-violet.Thi
- Page 76 and 77: 71* Salix alba. White willow.3 -+-t
- Page 78 and 79: 73Salix repens var. repens (var. er
- Page 80 and 81: 75*Rorippa sylvestris. Creeping yel
- Page 82 and 83: 77* Thlaspi arvense. Field pennycre
- Page 84 and 85: 79aphanus raphanistrum. Wild radish
- Page 86 and 87: ica tetralix. Cross~leaved heath.As
- Page 88 and 89: centre of the Forest on alluvial so
- Page 90 and 91: It is predominantly a plant which o
- Page 92 and 93: 87SAXIFRAGACEAE[*Darmera pe/tata. I
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89insipid in flavour IHaskel1 1960)
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91ection Rubus. Subsection Hiemales
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93Rubus pr%ngatus.Nutley (45.27), 1
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95bus vestitus.Newbridge, 1969, B.
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97Rubus largificus.3 -+t--------'+_
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Potentilla anserina. Silverweed.Com
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101Geum urbanum. Wood avens.Primari
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103osa x pseudorusticana fR. arvens
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105Prunus spinosa. Blackthorn, Wint
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orbus aucuparia. Rowan, Mountain as
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109r*Crataegus persimilis. Broad-le
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111icia tetrasperma. Smooth tare.Fo
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113Trifolium rep ens. White clover"
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115Genista tinctoria. Dyer's greenw
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*Genista anglica. Petty whin, Needl
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Myriophyllum alterniflorum. Alterna
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121Epilobium montanum x obscurum.Fo
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habitats.It lives up to the name of
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125,UPHORBIACEAEVlercurialis perenn
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Although one of Britain's tiniest p
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129*Oxalis articulata. Pink-sorrel.
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*Erodium cicutarium. Common stork's
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Widespread in Europe from the shore
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* Sison amomum. Stone parsley.Tetra
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137Centaurium pu/chellum. Lesser ce
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139CONVOLVULACEAEConvolvulus arvens
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141*Nymphoides pe/tata. Fringed wat
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Myosotis secunda (M. repens). Creep
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145Stachys 8lvensis. Field woundwor
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5cutellaria x hybrida (5. galericul
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Origanum vu/gare. Marjoram.Near Fis
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CALLlTRICHACEAE151Ca/litriche is a
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153Fraxinus excelsior. Ash.4 5Local
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~'Cymbalaria mural is. Ivy-leaved t
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157Veronica scutellata. Marsh speed
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159Sibthorpia europaea. Cornish mon
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161OROBANCHACEAEOrobanche rapum~gen
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163RUBIACEAESherardia arvensis. Mad
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165Viburnum OpUIU5.Guelder-rose, St
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167Valeriana officina/is 5.1. Commo
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169Cirsium vu/gate. Spear thistle.W
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171We suspect that Leontodon hispid
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173Mycelis muralis. Wall lettuce.Se
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175Taraxacum exceJlens.Scattered al
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177Taraxacum sinuatum.Recorded from
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179Crepis capillaris. Smooth hawk'
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181Hieracium. Hawkweeds.3134 5Hiera
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*So/idago gigantea. Early goldenrod
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ecorded with Chamaemelum are Matric
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*Senecio x albescens (S. cineraria
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*Petasites japonicus. Giant butterb
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*E1odea canadensis. Canadian waterw
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Arum maculatum. Lords~and~ladies, C
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Juncus x surrejanus (J. acutiflorus
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fences or topography. It may domina
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Eleocharis multicaulis. Many-stalke
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A hybrid first described by Col em
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203Carex riparia. Greater pond~sedg
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Material which initially looked lik
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207Festuca pratensis. Meadow fescue
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209Cynosurus cristatus. Crested dog
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211Glyceria f/uitans. Floating swee
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213Deschampsia cespitosa. Tufted ha
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Agrostis gigantea. Black bent.Tetra
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217Bromus lepidus. Slender soft~bro
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* Triticum aestivum. Bread wheat.A
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221L1L1ACEAENarthecium ossifragum.
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223*Allium triquetrum. Three-Gornar
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Trumpet length(mm)454035302520•
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227ORCHIDACEAEEpipactis purpurata.
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een unable to refind any bog orchid
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231DactylorhizB praetermissa. South
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Rachel A. Nicholson (1934-). Rachel
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235REFERENCESAbbas, M. H. (1979). S
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Graham, G. G. & Primavesi, A. L. (1
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Rich, T. C. G. & Smith, P. A. (1996
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GAZETTEER241This gazetteer has been
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Smockfarthing 40.31Smuggler's car p
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Asple adi+cuttr int~~~tl·Bteel SI!
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Broad bean 111 Cardamine x hausskne
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Fen bedstraw 163 Fumitory 46 Greate
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Leucanthemum maximum 186 Many-flowe
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Quercus ilex 50 Rorippa x ster/lis
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Sticky mouse-ear 56 Thale cress 73