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Flora of Ashdown Forest - Botanical Society of the British Isles

Flora of Ashdown Forest - Botanical Society of the British Isles

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Myriophyllum alterniflorum. Alternate water-milfoil.Pond near Coleman's Hatch, C. E. Salmon (Wolley-Dod 19371.One isolated record in <strong>the</strong> fishing lake at <strong>Forest</strong> Row (422.3471. 1995, TR, and <strong>the</strong>n aseries <strong>of</strong> ponds and streams along <strong>the</strong> Pippingford watercourse downstream from <strong>the</strong> pond inChelwood Vachery (431.2951. 1995, MR, PR & TR, to pond in Mill Wood (437.2881, 1995,MM & TR, to a pond (448.2991 and several lakes (44.301 in Pippingford Park, 1995, SBRSetc, In 1995 it flowered freely, possibly because <strong>of</strong> <strong>the</strong> warm wea<strong>the</strong>r.This was recorded in Hall (1980) from five tetrads just south-west <strong>of</strong> <strong>the</strong> <strong>Forest</strong> but noton it, and in Sussex is quite a rare plant. It tends to occur in base-poor and nutrient-poor4 5water such as drains from <strong>the</strong> acidic, uncultivated areas <strong>of</strong> <strong>the</strong> <strong>Forest</strong>, and is most frequent in<strong>the</strong> north and west <strong>of</strong> Britain. Mainly in west, north and central Europe, Greenland and <strong>the</strong> Azores.GUNNERACEAE[Gunnera manicata. Brazilian giant-rhubarb.Bottom <strong>of</strong> Dodd's Hill (450.2691, 1995, AK. Planted around pond at Chelwood Vachery (431.2971, 1994, TR, but notnaturalized. The species is spreading vegetatively in damp woodlands and meadows in a number <strong>of</strong> parts <strong>of</strong> Britain, andspreading by seed in <strong>the</strong> Channel Islands and western Ireland. It is native in sou<strong>the</strong>rn BrazilJLYTHRACEAELythrum salicaria. Purple loosestrife.Tetrads 43H and 43M along <strong>the</strong> Medway (Hall 19801.One plant in wood east <strong>of</strong> ford at Shalesbrook (435.3421. 1995, PW; damp ground onfield edge near Lines Farm (443.348), 1995, <strong>Flora</strong> meeting; several clumps in dryish ditch atbottom <strong>of</strong> Oldlands Hall formal gardens (476.2741. 1995, R. Barley et al.; Pound gate (48.281,3 1994, TR.Mainly in <strong>the</strong> main rivers and associated wetlands in Sussex (Hall 1980), and quitewidespread in <strong>the</strong> lowlands <strong>of</strong> Britain. It is decreasing in England (Rich & Woodruff 1996).Widespread in Europe except <strong>the</strong> far north, Asia and North Africa.4 5 This beautiful purplish-pink plant was introduced to North America in <strong>the</strong> 1800s, andnow billions <strong>of</strong> plants blanket many <strong>of</strong> <strong>the</strong>ir finest wetlands. Plants can produce 2.7 million seeds annually which aredispersed by water, animals and birds, and it is a severe threat to <strong>the</strong> natural flora and fauna as nothing <strong>the</strong>re eats it. Agood biological control agent would be worth patenting!Lythrum portula. Water-purslane.Considering that it was only recorded in tetrad 43L in Hall (1980) when it must have beenoverlooked, we have recorded it very widely. It is very characteristic <strong>of</strong> <strong>the</strong> wet hollows inmuddy tracks, and in shallow ponds with base-poor and nutrient-poor water. In <strong>the</strong> hotsummer <strong>of</strong> 1995 it was conspicuous turning reddish as it dried, but still we only added aproportional one third <strong>of</strong> records so not necessarily more abundant in dry years.Scattered throughout <strong>the</strong> Weald in wet places in open communities, and possibly <strong>the</strong> areain which it is most common in Britain. Widespread in Europe except <strong>the</strong> far north, Siberia and<strong>the</strong> Azores.This species shows a cline in <strong>the</strong> length <strong>of</strong> <strong>the</strong> calyx segments ~Allen 1954), with plantsin western Britain and Europe having <strong>the</strong>m 1.5-2 mm long (subsp. longidentata) compared to 0.5 mm in plants fromelsewhere (subsp. portula), though why is unknown. Plants from Jack Daw had very short calyx segments, but <strong>the</strong>y arelonger in o<strong>the</strong>r plants; we have not investigated <strong>the</strong>m in detail.THYMELAEACEAEDaphne laureola. Spurge-Iaurel.One plant on eastern edge <strong>of</strong> Courtland Wood (451.2611, 1995, TR; five plants by Marlpits(467.2641. 1993, TR & NM.The occurrence <strong>of</strong> this 'calcicole' at <strong>the</strong> Marlpits might be expected to be related tocalcareous clay, yet <strong>the</strong> pH <strong>of</strong> <strong>the</strong> clay was 3.9 and <strong>the</strong> leaf litter around was 4.6 (<strong>the</strong>se were3 -1---1-------1.--1- double-checkedl, and at Courtland Wood pH 5.3. At Marlpits <strong>the</strong> plants were also quitedamaged by grazing with many weak shoots, and <strong>the</strong>y flowered only very sparsely.4 5Mainly in woodlands on <strong>the</strong> chalk in Sussex, and rarer on <strong>the</strong> clays in <strong>the</strong> Weald (Hall1980). Locally common in England and Wales in chalk and limestone woodlands. It reachesits nor<strong>the</strong>rn-most distribution in Britain, and occurs in sou<strong>the</strong>rn and western Europe andtemperate Asia.119

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