91ection Rubus. Subsection Hiemales.eries Sylvatici.ubus chloophyllus fR. rhodanthus, R. rhombifoliusi.<strong>Ashdown</strong> <strong>Forest</strong>, 1899, T. Hilton IBM). Newbridge, 1969, B. A. Miles IPhilp 1970). Openwoods and heaths, particularly in <strong>Ashdown</strong> <strong>Forest</strong> (Edees & Newton 1988). The record for R.holerythrus from near Wych Cross, A. H. Wolley-Dod IWolley-Dod 1937) probably belongshere.3 --++------"+_ Common in woods near Pooh car park (472.332), 1994, DE, and possibly this species onheath opposite Townsend car park 1445.328), 1994, DE. Near Coldharbour 140.32), HindleapWarren 141.321. Coleman's Hatch 144.32), Gills Lap 146.31),1994, AN & AB.A characteristic species <strong>of</strong> <strong>Ashdown</strong> <strong>Forest</strong> in open woods and heaths. It also occurs in4 5Kent and western France.ubus imbricatus.TQf4.3 IHall 1980; Edees & Newton 1988).Nutley 144.27, 45.27, 45.28), Fairwarp 146.261. near Poundgate car park 1481.284),1995, RR & PH.Locally frequent in sou<strong>the</strong>rn Britain in well-defined populations on acidic soils onhedgebanks, heaths and borders <strong>of</strong> woods. Also known in France and Germany.* Rubus laciniatus.There are a number <strong>of</strong> scattered records from around <strong>the</strong> <strong>Forest</strong> for this introduced spec'leswhich has probably been spread from gardens by birds. Instantly distinguished from <strong>the</strong> o<strong>the</strong>rbrambles, even by <strong>the</strong> non-expert, by <strong>the</strong> finely divided leaflets,Widespread in north-western Europe.[Rubus lentiginosus.Newbridge, E. G. Gilbert (Wolley-Dod 1937). A very doubtful record as <strong>the</strong> main area <strong>of</strong> distribution is in North Wales, but<strong>the</strong>re are isolated records for <strong>the</strong> Hampshire-Berkshire border; it may refer to R. brevistaminosus which occurs atNewbridge.lRubus lindleianus.ilshdown <strong>Forest</strong>, ra<strong>the</strong>r scarce, H. J. Riddelsdell IWolleY-Dod 1937); Wych Cross, 1906, C. R. Billups IOXF, <strong>the</strong> TW4.3'ecord in Hall 1980; Edees & Newton 1988).Widespread in north-western Europe <strong>of</strong>ten on hedgebanks. The fruits <strong>of</strong> this species are ra<strong>the</strong>r small, seedy and sour:Earl 1994).~ubus platyacanthus (R. carpinifolius).'osingford Wood, W. C. Barton and H. J. Riddelsdell; <strong>Ashdown</strong> <strong>Forest</strong>, scarce, H. J. Riddelsdell IWolley-Dod 1937).:oleman's Hatch, W. C. Barton. TQ/4.3 IHall 1980; Edees & Newton 1988).Widespread in north-western Europe on heaths, etc.~ubus pyramidalis.Roadside between Newbridge and Chuck Hatch, 1934, W. C. Buton & H. J. Riddelsdell IBM).Stonehill 145.28), Gills Lap 146.311. roadsides near Suntinll's, Newbridge 146.32), 1994,AN & AB. Nutley 144.27, 45.27, 45.281. Fairwarp 146.23), near Poundgate car park1481.284), 1995, RR & PH.Widespread in north-western Europe.
92Rubus sciocharis.Gills Lap (46.31), 1994, AN & AB, <strong>the</strong> first record for East Sussex.Widespread in north-western Europe on wood borders and hedge banks but ra<strong>the</strong>rlocalised in Britain and possibly still colonising from <strong>the</strong> continent (RR, pers. comm. 1996).Series Rhamnif<strong>of</strong>li'.Rubus cardiophyllu5 fR. rhamnifolius}.Posingford Wood, E. G. Gilbert; fairly common on <strong>Ashdown</strong> <strong>Forest</strong>, H. J. Riddelsdell (Wolley·Dod 1937(. Newbridge, 1969, B. A. Miles (Philp 19701. TO/4.3 (Hall 1980; Edees & Newton19881.Roadsides near Sunting's, Newbridge (46.32), 1994, AN & AB. Nutley (45.27), nearEllison's Pond (46.28), Fairwarp Farm (47.26), Poundgate car park (48.28), 1995, RR & PH.Local in north-west Europe on edges <strong>of</strong> heaths, woods, and on hedgebanks butwidespread and common in most <strong>of</strong> lowland Britain.Rubus cissburiensis.Near Newbridge, 1934, W. C. Barton & H. J. Riddelsdell (BM(. TO/4.3 (Hall 1980; Edees &Newton 1988).Priory Road near Kidbrooke Wood (412.338), Goat car park (403.326), Warren car park(416.320), Townsend car park (445.328), 1994, DE. Near Coldharbour (40.32), Hindleap3 -++-------'+- Warren (41.32), Coleman's Hatch (44.32), Stonehill (45.28), Gills Lap (46.31), roadsides nearSunting's, Newbridge (46.32), east <strong>of</strong> Camp Hill (47.28), 1994, AN & AB. Nutley (44.27,45.27, 45.28), near Ellison's Pond (46.28), Fairwarp (46.26, 47.26), Toll Lane (461.263),near Poundgate car park (481.284), 1995, RR & PH.4 5Described from Cissbury in Sussex, and a regional endemic <strong>of</strong> heaths and wood bordersespecially near London. Widespread, <strong>of</strong>ten abundant in south-east England.Rubus dumnoniensis.Hindleap Warren (41.32), Stonehill (45.28), 1994, AN & AB (Edees & Newton 1988). Nutley(45.27), near Poundgate car park (481.284), 1995, RR & PH.Widespread in Britain, but a predominantly western species. Recently found in France.Rubus polyan<strong>the</strong>mus.3 -++----.-!.+_"4 5Wych Cross, A. H. Wolley·Dod; Posingford Wood, H. J. Riddelsdell (Wolley·Dod 1937).Marshy ground near Crowborough Warren, 1932, W. C. Barton (BM). Crowborough Common,1933, W. C. Barton & H. J. Riddelsdell (BM). TQ/4.3 (Hall 1980; Edees & Newton 1988).Warren car park (416.320), and Townsend's car park (445.328), 1994. DE. NearColdharbour (40.32), Hindleap Warren (41.32), Stone Hill (45.28), Gills Lap (46.311. roadsidesnear Sunting's, Newbridge (46.32), 1994, AN & AB. Nutley (44.27, 45.27), Fairwarp (46.26,47.26), near Ellison's Pond (46.28), 1995, RR & PH.Widespread in Britain in hedges, wood edges, quarries, etc. and in north-western Europe.
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Flora of Ashdown Forest,-,;I
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Copyright Tim Rich, Pat Donovan, Pa
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ACKNOWLEDGEMENTSWe would like to th
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THE HABITATS OF ASH DOWN FOREST3His
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Near the bottom of all the non~wood
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Conopodium majusCrataegus laevigata
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Fincham (1995) published a small ph
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13ANAL VS IS OF THE FLORA DATAIn th
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15The number of different visits to
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17Hedgerows Disturbed/waste ground
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19The frequencies with which specie
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21(r~0.434, pO.l).-0.059,•300t350
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23A second type of comparison has b
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25Similarly, environmental variable
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SPECIES ACCOUNTS27Lat'1n names foll
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29EQUISETACEAEEquisetum f/uviatile.
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One plant on a vertical, north-faci
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Viola pa/ustris in an open alder co
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35DRYOPTERIDACEAEPolystichum aculea
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37BLECHNACEAEBlechnum spicant. Hard
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39AngiospermsLAURACEAE* Laurus nobi
- Page 46 and 47: 41Ranunculus repens, Creeping butte
- Page 48 and 49: wet mud, small streams, ditches and
- Page 50 and 51: 45*Meconopsis cambrica. Welsh poppy
- Page 52 and 53: 47Ulmusx hollandica f?U. glabra x m
- Page 54 and 55: 49FAGACEAEFagus sylvatica. Beech.Fo
- Page 56 and 57: A special large oak at Chuck Hatch,
- Page 58 and 59: Cory/us avel/ana. Hazel.Recorded in
- Page 60 and 61: plants in the autumn; we have not r
- Page 62 and 63: Sagina procumbens. Procumbent pearl
- Page 64 and 65: 59*Saponaria officinalis. Soapwort,
- Page 66 and 67: 61Dolygonum arenastrum. Equal-leave
- Page 68 and 69: 63Rumex conglomeratus. Clustered do
- Page 70 and 71: 65Hypericum humifusum. Trailing S1
- Page 72 and 73: 67DROSERACEAEDrosera rotundifolia.
- Page 74 and 75: 69Viola laetea. Pale dog-violet.Thi
- Page 76 and 77: 71* Salix alba. White willow.3 -+-t
- Page 78 and 79: 73Salix repens var. repens (var. er
- Page 80 and 81: 75*Rorippa sylvestris. Creeping yel
- Page 82 and 83: 77* Thlaspi arvense. Field pennycre
- Page 84 and 85: 79aphanus raphanistrum. Wild radish
- Page 86 and 87: ica tetralix. Cross~leaved heath.As
- Page 88 and 89: centre of the Forest on alluvial so
- Page 90 and 91: It is predominantly a plant which o
- Page 92 and 93: 87SAXIFRAGACEAE[*Darmera pe/tata. I
- Page 94 and 95: 89insipid in flavour IHaskel1 1960)
- Page 98 and 99: 93Rubus pr%ngatus.Nutley (45.27), 1
- Page 100 and 101: 95bus vestitus.Newbridge, 1969, B.
- Page 102 and 103: 97Rubus largificus.3 -+t--------'+_
- Page 104 and 105: Potentilla anserina. Silverweed.Com
- Page 106 and 107: 101Geum urbanum. Wood avens.Primari
- Page 108 and 109: 103osa x pseudorusticana fR. arvens
- Page 110 and 111: 105Prunus spinosa. Blackthorn, Wint
- Page 112 and 113: orbus aucuparia. Rowan, Mountain as
- Page 114 and 115: 109r*Crataegus persimilis. Broad-le
- Page 116 and 117: 111icia tetrasperma. Smooth tare.Fo
- Page 118 and 119: 113Trifolium rep ens. White clover"
- Page 120 and 121: 115Genista tinctoria. Dyer's greenw
- Page 122 and 123: *Genista anglica. Petty whin, Needl
- Page 124 and 125: Myriophyllum alterniflorum. Alterna
- Page 126 and 127: 121Epilobium montanum x obscurum.Fo
- Page 128 and 129: habitats.It lives up to the name of
- Page 130 and 131: 125,UPHORBIACEAEVlercurialis perenn
- Page 132 and 133: Although one of Britain's tiniest p
- Page 134 and 135: 129*Oxalis articulata. Pink-sorrel.
- Page 136 and 137: *Erodium cicutarium. Common stork's
- Page 138 and 139: Widespread in Europe from the shore
- Page 140 and 141: * Sison amomum. Stone parsley.Tetra
- Page 142 and 143: 137Centaurium pu/chellum. Lesser ce
- Page 144 and 145: 139CONVOLVULACEAEConvolvulus arvens
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141*Nymphoides pe/tata. Fringed wat
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Myosotis secunda (M. repens). Creep
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145Stachys 8lvensis. Field woundwor
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5cutellaria x hybrida (5. galericul
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Origanum vu/gare. Marjoram.Near Fis
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CALLlTRICHACEAE151Ca/litriche is a
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153Fraxinus excelsior. Ash.4 5Local
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~'Cymbalaria mural is. Ivy-leaved t
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157Veronica scutellata. Marsh speed
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159Sibthorpia europaea. Cornish mon
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161OROBANCHACEAEOrobanche rapum~gen
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163RUBIACEAESherardia arvensis. Mad
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165Viburnum OpUIU5.Guelder-rose, St
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167Valeriana officina/is 5.1. Commo
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169Cirsium vu/gate. Spear thistle.W
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171We suspect that Leontodon hispid
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173Mycelis muralis. Wall lettuce.Se
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175Taraxacum exceJlens.Scattered al
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177Taraxacum sinuatum.Recorded from
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179Crepis capillaris. Smooth hawk'
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181Hieracium. Hawkweeds.3134 5Hiera
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*So/idago gigantea. Early goldenrod
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ecorded with Chamaemelum are Matric
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*Senecio x albescens (S. cineraria
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*Petasites japonicus. Giant butterb
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*E1odea canadensis. Canadian waterw
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Arum maculatum. Lords~and~ladies, C
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Juncus x surrejanus (J. acutiflorus
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fences or topography. It may domina
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Eleocharis multicaulis. Many-stalke
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A hybrid first described by Col em
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203Carex riparia. Greater pond~sedg
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Material which initially looked lik
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207Festuca pratensis. Meadow fescue
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209Cynosurus cristatus. Crested dog
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211Glyceria f/uitans. Floating swee
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213Deschampsia cespitosa. Tufted ha
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Agrostis gigantea. Black bent.Tetra
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217Bromus lepidus. Slender soft~bro
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* Triticum aestivum. Bread wheat.A
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221L1L1ACEAENarthecium ossifragum.
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223*Allium triquetrum. Three-Gornar
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Trumpet length(mm)454035302520•
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227ORCHIDACEAEEpipactis purpurata.
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een unable to refind any bog orchid
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231DactylorhizB praetermissa. South
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Rachel A. Nicholson (1934-). Rachel
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235REFERENCESAbbas, M. H. (1979). S
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Graham, G. G. & Primavesi, A. L. (1
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Rich, T. C. G. & Smith, P. A. (1996
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GAZETTEER241This gazetteer has been
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Smockfarthing 40.31Smuggler's car p
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Asple adi+cuttr int~~~tl·Bteel SI!
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Broad bean 111 Cardamine x hausskne
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Fen bedstraw 163 Fumitory 46 Greate
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Leucanthemum maximum 186 Many-flowe
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Quercus ilex 50 Rorippa x ster/lis
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Sticky mouse-ear 56 Thale cress 73