111icia tetrasperma. Smooth tare.Four tetrads on <strong>the</strong> <strong>Forest</strong> (Hall 1980).Usually found as a few plants scattered along rides, but never common, and in <strong>the</strong>farmland around. Twyford area (39.31), 1993/4, DB; south <strong>of</strong> Nutley (44.261, 1993, MM;near lake, Lines Farm (44.34),1995, PW; Newbridge (457.326),1994, <strong>Flora</strong> meeting; 46.31,3 -+t-----'-t- 1993, PO; south <strong>of</strong> Chuck Hatch (473.329),1994, TR; 49.31,1993, MM.Hall (1980) noted that in Sussex it prefers damper ground than V. hirsuta, and this mayexplain why it is more frequent on <strong>the</strong> silty <strong>Forest</strong> soils. It is widespread in south-eastEngland, and rarer elsewhere in Britain. Widespread in Europe except <strong>the</strong> far north; western4 5Asia, Japan and North Africa.Vicia sepium. Bush vetch.3 -+.----""4 5Many tetrads on <strong>the</strong> <strong>Forest</strong>, but one <strong>of</strong> <strong>the</strong> few places it is not ubiquitous in <strong>the</strong> Sussex Weald(Hall 1980).Scattered around <strong>the</strong> <strong>Forest</strong> in <strong>the</strong> villages and on roadsides and hedges, and occasionallyin open woodland. It is quite characteristic <strong>of</strong> <strong>the</strong> bushy edges <strong>of</strong> car parks.In south-east England it tends to occur on moist soils <strong>of</strong>ten in partial shade but is lessrestricted in <strong>the</strong> north and west, perhaps reflecting a requirement for moisture (Grime et al.1988). Widespread in Britain and Europe, though rare in <strong>the</strong> Mediterranean region, temperateAsia and Greenland.Vicia sativa. Common vetch, Fitches.This is an aggregate with a complex taxonomic history which means <strong>the</strong> old records aredifficult to interpret, and should be treated as Vicia sativa sensu lata (e.g. Hall 1980, etc.).Like V. sepium it is scattered around <strong>the</strong> <strong>Forest</strong> on roadsides and in <strong>the</strong> villages. Thecommonest taxon <strong>of</strong> this group on <strong>the</strong> <strong>Forest</strong> is subsp. segetalis and probably all records canbe referred to this. Very narrow-leaved plants were noted on track sides south <strong>of</strong> Brown'sBrook (472.2771, 1994, <strong>Flora</strong> meeting and could be subsp. nigra.Widespread in Britain, and in Europe except <strong>the</strong> far north; temperate Asia and North4 5Africa.* Vicia !aba. Broad bean.One plant on verge to Hunters Farm (449.2651, possibly carried by pigeons from nearby field, 1995, TR & G. M. Kay.Gone by January 1996, probably eaten by a hungry bunny.Lathyrus linifolius (L. montanus). Bitter-vetch.Common on <strong>the</strong> <strong>Forest</strong> (Hall 1980).Most typical <strong>of</strong> hedge banks and verges, sometimes in sun or light shade, usually onmoderately but not strongly acidic soils. A lovely plant and one <strong>of</strong> TR's favourites.Widespread in <strong>the</strong> Weald in Sussex but rare elsewhere. It occurs throughout Britain in4 5woodland, moorland, unimproved grassland and hedgebanks on acidic to neutral soils; many <strong>of</strong><strong>the</strong> sites are somewhat damp or humid. It is decreasing in England, presumably due to habitatloss (Rich & Woodruff 1996). Widespread in west, central and sou<strong>the</strong>rn Europe to Russia, andin North Africa.*Lathyrus latifolius. Broad-leaved everlasting-pea.Firebreak on heath imported on soil used to improve <strong>the</strong> track, Duddleswell (465.282), 1993,PW.A garden escape or throw-out now widely established and 'Increasing 'In Britain onroadsides, railway banks and waste ground {Rich & Woodruff 1996). Native in sou<strong>the</strong>rn and3 -++_-----'-+_ central Europe and North Africa.4 5
112Lathyrus nissolia. Grass vetchling, Shoes-and-stockings.,--,---------,------, Several plants on dumped soil outside Twyford Farm (397.312), 1994, DB & TR, but where<strong>the</strong> soi! came from is ano<strong>the</strong>r matter! This species can be very difficult to see amongst grassias its leaves match <strong>the</strong> grass leaves, but <strong>the</strong> red flowers or <strong>the</strong> pale brown pods usually give itiaway.IPredominantly recorded in south-east England and frequent in Sussex, It seems to belincreasing by spreading along major road verges (e.g. along <strong>the</strong> M4; Rich 1994c), a favouredlhabitat being <strong>the</strong> tall Arrhena<strong>the</strong>rum grassland. In Sussex it has a patchy distributionapparently unrelated to soils (Hall 1980), Widespread in west, central and sou<strong>the</strong>rn Europe to45<strong>the</strong> Caucasus and Syria; North Africa.Lathyrus pratensis. Meadow vetchling, Tom thumb.,-,----===,-----,----, Frequent in meadows where not heavily grazed and on verges around <strong>the</strong> <strong>Forest</strong> and mostiabundant on <strong>the</strong> clays or in limed meadows where <strong>the</strong> soil pH is slightly higher. The map!]shows it occurs mainly around <strong>the</strong> edges <strong>of</strong> <strong>the</strong> <strong>Forest</strong>, and it is absent from <strong>the</strong> heathland andwoodland. :Ubiquitous in Sussex (Hall 1980), in Britain and widespread in Europe. It is also found inlAsia from Siberia south to <strong>the</strong> Himalaya, and in North Africa.i*Me/i/otus a/bus. White melilot.Tetrad 42Z (Hall 1980), not refound.Usually a casual scattered in Sussex on disturbed ground, and more established in <strong>the</strong> larger conurbations. Native inEurasia and North Africa.Medicago lupulina.4Black medick, Nonesuch.Our maps shows some interesting lines <strong>of</strong> records mainly related to <strong>the</strong> roads - it tends tooccur on <strong>the</strong> more base-rich soils in open grassland on <strong>the</strong> verges which agrees with <strong>the</strong>observations <strong>of</strong> Grime et al. (1988) who note it is commonest on calcareous, moderately,disturbed infertile soils in Britain and is essentially a colonist <strong>of</strong> bare areas. Calcareous dustfrom limestone chippings used to surface <strong>the</strong> roads gets deposited on <strong>the</strong> verges.Recorded in Sussex in 90% <strong>of</strong> <strong>the</strong> tetrads (Hall 1980), and in lowland Britain.Widespread in Europe except <strong>the</strong> far north, temperate Asia and North Africa. Sometimescultivated for forage.5Medicago sativa subsp. sativa. Lucerne.A few plants in rough grass at <strong>the</strong> sou<strong>the</strong>rn end <strong>of</strong> Duddleswell car park (466.273), 1995, TR;Newnham area (49.28), 1994, SBRS.Rarely grown now as a fodder crop though it was formerly commonly so, andconsequently decreasing as an escape from cultivation (Rich & Woodruff 1996). The plants at3 Duddleswell may have originated from seed used to sow <strong>the</strong> car park, and it is also sometimesfound in bird seed. Locally frequent on <strong>the</strong> chalk in Sussex, and scattered through lowlandEngland. Native in Eurasia and North Africa.4 5Medicago arabica. Spotted medick.Bank <strong>of</strong> Long car park on foreign soil (427.311), 1995, PW; Chelwood Vachery (43.30),1994, <strong>Flora</strong> meeting; dumped soil in Pippingford Park (444.303), 1993, SBRS; verge at TollLane (462.263), 1993, TR.Very much a warmth-loving plant, and scattered along <strong>the</strong> coastal region in Sussex with3 -++--'"'''"-----'__\_ fewer records in <strong>the</strong> Weald. Similarly in Britain it is only frequent in <strong>the</strong> extreme south-east <strong>of</strong>England, and occurs scattered around <strong>the</strong> coast elsewhere. Mainly in sou<strong>the</strong>rn Europe,4 5extending north to Britain and <strong>the</strong> Ne<strong>the</strong>rlands, and in North Africa. ,II
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Flora of Ashdown Forest,-,;I
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Copyright Tim Rich, Pat Donovan, Pa
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ACKNOWLEDGEMENTSWe would like to th
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THE HABITATS OF ASH DOWN FOREST3His
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Near the bottom of all the non~wood
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Conopodium majusCrataegus laevigata
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Fincham (1995) published a small ph
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13ANAL VS IS OF THE FLORA DATAIn th
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15The number of different visits to
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17Hedgerows Disturbed/waste ground
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19The frequencies with which specie
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21(r~0.434, pO.l).-0.059,•300t350
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23A second type of comparison has b
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25Similarly, environmental variable
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SPECIES ACCOUNTS27Lat'1n names foll
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29EQUISETACEAEEquisetum f/uviatile.
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One plant on a vertical, north-faci
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Viola pa/ustris in an open alder co
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35DRYOPTERIDACEAEPolystichum aculea
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37BLECHNACEAEBlechnum spicant. Hard
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39AngiospermsLAURACEAE* Laurus nobi
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41Ranunculus repens, Creeping butte
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wet mud, small streams, ditches and
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45*Meconopsis cambrica. Welsh poppy
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47Ulmusx hollandica f?U. glabra x m
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49FAGACEAEFagus sylvatica. Beech.Fo
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A special large oak at Chuck Hatch,
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Cory/us avel/ana. Hazel.Recorded in
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plants in the autumn; we have not r
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Sagina procumbens. Procumbent pearl
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59*Saponaria officinalis. Soapwort,
- Page 66 and 67: 61Dolygonum arenastrum. Equal-leave
- Page 68 and 69: 63Rumex conglomeratus. Clustered do
- Page 70 and 71: 65Hypericum humifusum. Trailing S1
- Page 72 and 73: 67DROSERACEAEDrosera rotundifolia.
- Page 74 and 75: 69Viola laetea. Pale dog-violet.Thi
- Page 76 and 77: 71* Salix alba. White willow.3 -+-t
- Page 78 and 79: 73Salix repens var. repens (var. er
- Page 80 and 81: 75*Rorippa sylvestris. Creeping yel
- Page 82 and 83: 77* Thlaspi arvense. Field pennycre
- Page 84 and 85: 79aphanus raphanistrum. Wild radish
- Page 86 and 87: ica tetralix. Cross~leaved heath.As
- Page 88 and 89: centre of the Forest on alluvial so
- Page 90 and 91: It is predominantly a plant which o
- Page 92 and 93: 87SAXIFRAGACEAE[*Darmera pe/tata. I
- Page 94 and 95: 89insipid in flavour IHaskel1 1960)
- Page 96 and 97: 91ection Rubus. Subsection Hiemales
- Page 98 and 99: 93Rubus pr%ngatus.Nutley (45.27), 1
- Page 100 and 101: 95bus vestitus.Newbridge, 1969, B.
- Page 102 and 103: 97Rubus largificus.3 -+t--------'+_
- Page 104 and 105: Potentilla anserina. Silverweed.Com
- Page 106 and 107: 101Geum urbanum. Wood avens.Primari
- Page 108 and 109: 103osa x pseudorusticana fR. arvens
- Page 110 and 111: 105Prunus spinosa. Blackthorn, Wint
- Page 112 and 113: orbus aucuparia. Rowan, Mountain as
- Page 114 and 115: 109r*Crataegus persimilis. Broad-le
- Page 118 and 119: 113Trifolium rep ens. White clover"
- Page 120 and 121: 115Genista tinctoria. Dyer's greenw
- Page 122 and 123: *Genista anglica. Petty whin, Needl
- Page 124 and 125: Myriophyllum alterniflorum. Alterna
- Page 126 and 127: 121Epilobium montanum x obscurum.Fo
- Page 128 and 129: habitats.It lives up to the name of
- Page 130 and 131: 125,UPHORBIACEAEVlercurialis perenn
- Page 132 and 133: Although one of Britain's tiniest p
- Page 134 and 135: 129*Oxalis articulata. Pink-sorrel.
- Page 136 and 137: *Erodium cicutarium. Common stork's
- Page 138 and 139: Widespread in Europe from the shore
- Page 140 and 141: * Sison amomum. Stone parsley.Tetra
- Page 142 and 143: 137Centaurium pu/chellum. Lesser ce
- Page 144 and 145: 139CONVOLVULACEAEConvolvulus arvens
- Page 146 and 147: 141*Nymphoides pe/tata. Fringed wat
- Page 148 and 149: Myosotis secunda (M. repens). Creep
- Page 150 and 151: 145Stachys 8lvensis. Field woundwor
- Page 152 and 153: 5cutellaria x hybrida (5. galericul
- Page 154 and 155: Origanum vu/gare. Marjoram.Near Fis
- Page 156 and 157: CALLlTRICHACEAE151Ca/litriche is a
- Page 158 and 159: 153Fraxinus excelsior. Ash.4 5Local
- Page 160 and 161: ~'Cymbalaria mural is. Ivy-leaved t
- Page 162 and 163: 157Veronica scutellata. Marsh speed
- Page 164 and 165: 159Sibthorpia europaea. Cornish mon
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161OROBANCHACEAEOrobanche rapum~gen
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163RUBIACEAESherardia arvensis. Mad
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165Viburnum OpUIU5.Guelder-rose, St
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167Valeriana officina/is 5.1. Commo
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169Cirsium vu/gate. Spear thistle.W
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171We suspect that Leontodon hispid
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173Mycelis muralis. Wall lettuce.Se
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175Taraxacum exceJlens.Scattered al
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177Taraxacum sinuatum.Recorded from
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179Crepis capillaris. Smooth hawk'
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181Hieracium. Hawkweeds.3134 5Hiera
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*So/idago gigantea. Early goldenrod
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ecorded with Chamaemelum are Matric
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*Senecio x albescens (S. cineraria
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*Petasites japonicus. Giant butterb
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*E1odea canadensis. Canadian waterw
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Arum maculatum. Lords~and~ladies, C
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Juncus x surrejanus (J. acutiflorus
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fences or topography. It may domina
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Eleocharis multicaulis. Many-stalke
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A hybrid first described by Col em
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203Carex riparia. Greater pond~sedg
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Material which initially looked lik
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207Festuca pratensis. Meadow fescue
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209Cynosurus cristatus. Crested dog
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211Glyceria f/uitans. Floating swee
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213Deschampsia cespitosa. Tufted ha
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Agrostis gigantea. Black bent.Tetra
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217Bromus lepidus. Slender soft~bro
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* Triticum aestivum. Bread wheat.A
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221L1L1ACEAENarthecium ossifragum.
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223*Allium triquetrum. Three-Gornar
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Trumpet length(mm)454035302520•
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227ORCHIDACEAEEpipactis purpurata.
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een unable to refind any bog orchid
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231DactylorhizB praetermissa. South
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Rachel A. Nicholson (1934-). Rachel
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235REFERENCESAbbas, M. H. (1979). S
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Graham, G. G. & Primavesi, A. L. (1
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Rich, T. C. G. & Smith, P. A. (1996
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GAZETTEER241This gazetteer has been
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Smockfarthing 40.31Smuggler's car p
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Asple adi+cuttr int~~~tl·Bteel SI!
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Broad bean 111 Cardamine x hausskne
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Fen bedstraw 163 Fumitory 46 Greate
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Leucanthemum maximum 186 Many-flowe
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Quercus ilex 50 Rorippa x ster/lis
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Sticky mouse-ear 56 Thale cress 73