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Flora of Ashdown Forest - Botanical Society of the British Isles

Flora of Ashdown Forest - Botanical Society of the British Isles

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171We suspect that Leontodon hispidus has been over-recorded for this species; Hypochaeris can be easily distinguishedin that it has simple not forked hairs on <strong>the</strong> leaves, and has pale yellow, triangular scales on <strong>the</strong> receptacle - this can beseen by plucking <strong>the</strong> florets out to leave <strong>the</strong> scales behind.Recorded in 92% <strong>of</strong> <strong>the</strong> Sussex tetrads (Hall 1980). Widely distributed throughout Britain, mainly on neutral tomoderately acidic, well-drained soils in grazed or mown pastures and on verges. Widespread in Europe, nor<strong>the</strong>rn Asia andNorth Africa.Leontodon autumnalis subsp. autumns/is. Autumn hawkbit.Common on <strong>the</strong> <strong>Forest</strong> (Hall 1980).Widespread and locally frequent on road verges, pastures, rides and waste places on wetor acidic soils. It flowers in late summer and is a very variable species, some plants havingalmost entire leaves and o<strong>the</strong>rs almost pinnate.It is easily distinguished from <strong>the</strong> o<strong>the</strong>r Leontodon species by <strong>the</strong> branched floweringstems and has only simple hairs on <strong>the</strong> leaves. It is separated from Hypochaeris by <strong>the</strong>absence <strong>of</strong> receptacular scales.Very common in Sussex and Britain. Widespread in Europe though rarer in <strong>the</strong> south. InAsia, North~west Africa and Greenland.Leontodon hispidus. Rough hawkbit.Recorded in ten tetrads in Hall (1980), but surely over~recorded, perhaps for Hypochaeris?Poundgate road verge where chalk may have been used to build <strong>the</strong> road (484.285), pH7.7, 1993 +, <strong>Flora</strong> meeting; Rystwood Road on clay (441.344), pH 6.6, 1995, TR; Nutleychurchyard, pH 6.6, Coleman's Hatch churchyard, pH 6.5, and St John's churchyard, pH 6.S,3 1995, PD.The rarity with which we have found this species compared to <strong>the</strong> apparent frequency in<strong>the</strong> Sussex Plant Atlas is surprising, and only two <strong>of</strong> <strong>the</strong> tetrads are in common. It is mostabundant on chalk and limestone soils <strong>of</strong> mid to high pH, and would not be expected to be4 5common on <strong>the</strong> <strong>Forest</strong> except in places with raised pH such as road verges as, ourmeasurements indicate. It is also characteristic <strong>of</strong> <strong>the</strong> churchyards; raised pH in soils around churches may be a result <strong>of</strong>calcium leaching from mortar ei<strong>the</strong>r directly from <strong>the</strong> buildings or from discarded material when <strong>the</strong>y are rebuilt, or perhapsfrom <strong>the</strong> tombstones. L. hispidus could have been present around Coleman's Hatch, where <strong>the</strong> soils are base~rich clay, andat Nutley, but must have been introduced elsewhere.It is similar to L. saxatliis in having forked hairs on <strong>the</strong> leaves, a single stem and drooping buds, but all <strong>the</strong> acheneshave a pappus, and <strong>the</strong> involucre is darker in colour and it is much more hairy (though L. saxatilis can sometimes be veryhairy too). Hybrids between <strong>the</strong> two have been recorded in Britain but we have not found any.Widespread in lowland Britain on calcareous soils and clays in unimproved pastures. Widespread in Europe and AsiaMinor.Leontodon saxatilis (L. taraxacoides). Lesser hawkbit.Locally frequent on <strong>the</strong> east side <strong>of</strong> <strong>the</strong> <strong>Forest</strong> (Hall 1980).Although it is common on <strong>the</strong> <strong>Forest</strong> on road verges, tracks, lawns and open, drygrassland it is very <strong>of</strong>ten overlooked as an intensive search in East Sussex by PO proved,resulting in a large increase in records (Briggs 1990).It is easily identified when in flower by <strong>the</strong> outer achenes which have no pappus, whilst<strong>the</strong> inner ones do. These outer fruits persist long after <strong>the</strong> inner ones have been blown awayand <strong>the</strong> stem is dead (even as late as November).Widespread in Sussex on heaths, on bare, open, generally acidic or sandy soils.Ubiquitous in Britain. Widespread in sou<strong>the</strong>rn, central and western Europe.Picris echioides. Bristly oxtongue.Scattered around <strong>the</strong> <strong>Forest</strong> on dumped soil, road verges and in car parks, but usually only afew plants at a time, and not recorded in Hall (1980).Common on <strong>the</strong> chalk and on clays, especially on roadsides and near <strong>the</strong> coast in Sussex.Locally abundant in south~easte.rn Britain and widespread in sou<strong>the</strong>rn Europe and South~west3 Asia.4 5

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